Revision of the genus Delenda Croissandeau, 1891 (Coleoptera: Staphylinidae: Pselaphinae)
Author
Bekchiev, Rostislav
Author
Brachat, Volker
text
Zootaxa
2014
3765
6
583
592
journal article
46380
10.11646/zootaxa.3765.6.6
1cc7e809-f704-41c0-a0b2-8f9608dc4ffd
1175-5326
228037
E9220330-C692-47A3-BADE-364BF2FB74D2
Delenda
Croissandeau, 1891
Delenda
Croissandeau, 1891
.
Type
species:
Delenda carthago
Croissandeau, 1891
, monotypy.
Delenda
Croissandeau
:
Raffray, 1904
: 500
(synonymy of
Eusonoma
Reitter
);
Raffray, 1911
: 7
(catalogue);
Newton & Chandler, 1989
: 18
(catalogue).
Eusonoma
Reitter, 1893
: 172
.
Type
species:
Eusonoma frivaldszkyi
Reitter, 1893
, monotypy.
Redescription.
Habitus as in
Fig. 1
; length
1.5
–
1.7 mm
; body relatively flat, covered with short, suberect golden setae. Head (
Fig. 2
a) wider than long (0.28
–
0.34 /
0.20
–
0.28mm
) with two deep vertexal foveae; frontal sulcus large and triangular, opened on frontal rostrum, which is slightly extended and rounded anteriorly; antennal tubercules small. Antennae (
Fig. 2
b) with eleven antennomeres; scape longer than wide (0.09
–
0.1/
0.048
–
0.05 mm
); pedicel longer than wide (0.06/
0.05 mm
); III as wide as long (0.03/
0.03 mm
); IV longer than wide (0.04/
0.03 mm
); antennomeres V–VII equal in size, wider than long (0.04/
0.04 mm
); antennal club 3-segmented, welldefined. Each eye composed of about 10
–
30 facets. Maxillary palpi (
Fig. 2
c) with palpomeres I short, II lengthily pedunculate, III small, IV large and oval. Labrum (
Fig. 3
) species characteristic; with few setae. Pronotum (
Fig. 4
a) wider than long (0.32
–
0.38 /
0.24
–
0.30 mm
), widest part situated anterior to middle; pronotal disc at middle with large flat area, with five elongate, shallow depressions, two in anterior part, and three in posterior part; lateral antebasal foveae deep, not connected by antebasal sulcus; basolateral foveae present, and connected by a fine sulcus. Prosternum (
Fig. 4
b) with median and lateral procoxal foveae. Mesoventrite (
Fig. 4
c) with median mesoventral fovea, and deep lateral mesoventral foveae, with lateral mesocoxal foveae. Metaventrite with lateral metaventral foveae. Elytra (
Fig. 5
a) very short, length
0.22
–
0.26 mm
, with width at base (
0.34
–
0.48 mm
); each elytron lacking basal foveae, with two discal foveae; sutural stria incomplete; without discal stria. Abdomen (
Fig. 5
b, c) length
0.93
–
1.02 mm
; first three visible tergites (IV–VI) wider than long (0.44
–
0.50/
0.18
–
0.20 mm
), without any trace of foveae or carinae; first three visible sternites (IV–VI, each with two basolateral foveae. Legs short and robust, metacoxae (
Fig. 5
d) with deep fovea on external border. All known species are wingless. Aedegus (
Fig. 6
) with median lobe thin and asymmetrical, parameres well-developed and symmetrical, located on large tooth of basal part of median lobe.
Sexual dimorphism.
Not apparent. The penial plate (sternite IX) of the male (
Fig. 5
b) is oval and long, while the female lacks this modification (
Fig. 5
c).
Biology. S
pecies of the genus are always collected by sifting leaf-litter in mesophilous
Fagus sylvatica
L. forests, with
Rhododendron ponticum
L. or
Platanus orientalis
Mill.
(
Fig. 7
a) present, and from xerophilous forests of
Quercus
spp., and
Caprinus orientalis
Mill.
, (
Fig. 7
b).
Distribution.
Bulgaria
,
Turkey
(
Fig. 8
)
Relationships.
Delenda
is a very peculiar genus that can be readily separated from other genera of Faronitae by the following combination of characters: head with large triangular sulcus; antennae with well-formed terminal club; elytra very short, with only two discal foveae, sutural stria incomplete; metacoxa with fovea. A relationship with
Faronus
Aub, 1844
could be possible, since both genera share a fovea on the metacoxae, but this is the only shared, excluding the common characters typical for Faronitae, character between them. These fovea are missing in some other genera of Faronitae such as
Sonoma
Casey, 1886
,
Golasa
Raffray, 1904
and
Golasina
Jeannel, 1962
(Kurbatov pers. comm). Aside from this possible affinity to
Faronus
,
Delenda
appears to be an isolated genus in the supertribe without clear relationships to any other known genus of Faronitae.