Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny Author R. G. Beutel Author S. N. Gorb text J. Zool. Syst. Evol. Research 2001 39 177 207 journal article 10.1046/j.1439-0469.2001.00155.x 95b3d62f-8fcc-4088-873f-b33f4a63d3fc 2360323 24 Heteroptera The tarsus is maximally three­segmented. An arolium is present in slaterellines ( Lygaeidae ), an arolium and smooth euplantulae in mirines ( Miridae ), smooth pulvilli are present in members of Coreidae , Pentatomidae and Pyrrhocoridae ( Figs 3b , 5A ), and smooth pulvilli and a fossula spongiosa in reduviids ( Carver et al. 1991 ) ( Fig. 3g ).