A taxonomic revision of two genera, Pleurolobus and Sohmaea (Leguminosae) in Thailand and Indo-China
Author
Saisorn, Witsanu
School of Science, Walailak University, Nakhon Si Thammarat 80161, Thailand.
Author
Chantaranothai, Pranom
Department of Biology & Centre of Excellence on Biodiversity (BDC), Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand.
text
Phytotaxa
2022
2022-11-23
573
2
231
246
http://dx.doi.org/10.11646/phytotaxa.573.2.4
journal article
194617
10.11646/phytotaxa.573.2.4
80d52cdf-fbd6-4aa4-b9f1-c6e8ac9dc473
1179-3163
7349951
2.
Sohmaea hispida
(
Franchet 1890: 174
) H.Ohashi & K.Ohashi (2018b: 162)
.
Fig. 1C–D
.
≡
Desmodium hispidum
Franchet
.
Type:—
CHINA
.
Yunnan
,
Delavay
3504
(
holotype
P
!
P02142548
, isotype A! A00214959 [digital image]).
≡
Uraria hispida
(Franch.)
Schindler (1926: 254)
.
=
Uraria henryi
Schindler (1925: 15)
. Type:—
CHINA
.
Yunnan
, Mengtze,
Henry 9342
(
lectotype
A!A00063421 [digital image], designated by
Ohashi
et al.
, 2002
).
Small shrub,
0.3–1 m
high; stems and twigs angular, deeply sulcate, densely silky and spreading uncinate hairy.
Leaves
trifoliolate, spirally arranged; stipules triangular to ovate, 8–13 ×
3–5 mm
, apex long acuminate, surface densely silky and uncinate hairy; petioles
1–2.5 cm
long, densely silky hairy; rachis
1–2 mm
long, densely silky hairy.
Leaflets
: stipels triangular to narrowly triangular, 5–7 ×
0.5–1 mm
, apex acuminate to long acuminate, surface densely hairy; petiolules
1–2 mm
long, densely hairy.
Terminal leaflet
broadly lanceolate to ovate, 5–14 ×
2–7 cm
, apex narrowly acuminate, base obtuse to rounded, margin entire, upper surface sparsely appressed pubescent and densely uncinate hairy, lower surface soft and densely ascending hairy, without uncinate hairs; lateral veins 9–13 per side, conspicuous.
Lateral leaflets
obliquely lanceolate to ovate, 3.5–10 ×
1.5–5 cm
, apex narrowly acuminate, base obliquely obtuse to rounded, margin entire, both surfaces like terminal leaflet; lateral veins 8–13 per side, conspicuous.
Inflorescences
simple, occasionally paniculate,
3–25 cm
long, terminal or axillary; rachis soft, densely and ascending pubescent.
Primary bract
narrowly triangular to ovate, 4–6 ×
1–1.5 mm
, apex narrowly acuminate, margin fimbriate, surface soft and densely hairy, enclosing 2-immature flowers and secondary bract.
Secondary bract
narrowly triangular to linear, 1.5–3 ×
0.2–0.4 mm
, pubescent.
Flowers
ca.
4 mm
long, borne in 2-flowered fascicles; bracteoles absent; pedicels
4–4.5 mm
long, with densely uncinate hairs.
Calyx
pale green, ca.
4 mm
long, base cuneate; outside long pubescent and spreading uncinate hairy, inside glabrous, tube ca.
1.2 mm
long; teeth
2–3 mm
long, lower one longer than others, upper tooth entire at the apex.
Corolla
: standard violet to white, wings and keels pale violet; standard obovate, ca. 4 ×
2.4 mm
, apex shallowly emarginated to rounded, base attenuate, not auriculate, claw ca.
0.1 mm
long; wings narrowly oblong, ca. 2.5 ×
1 mm
, apex obtuse, base auriculate, claw ca.
0.8 mm
long; keels narrowly oblong, curved, ca. 4 ×
1 mm
, apex obtuse, claw ca.
0.8 mm
long.
Stamens
ca.
3 mm
long.
Gynoecium
: ovary uncinate hairy, 4–6-ovulate.
Pods
brown, sessile, plicate when young, becoming straight when mature, indehiscent, 6–7-articulate, oblong,
1.8–2.1 cm
long,
1.8–2 mm
wide, surface densely uncinate hairy, striate reticulate veined, both upper and lower sutures less constricted; isthmus 1.5/2 as broad as the pod; articles oblong, ca.
3 mm
long; fruiting pedicels
4.5–5 mm
long, reflexed, with densely uncinate hairs.
Seeds
dark brown, transversely oblong, ca. 2 × 01 mm, ca.
0.5 mm
thick, arillate around hilum, the area round hilum not concave.
Distribution
:—
India
,
Myanmar
,
China
, and
Thailand
.
Ecology
:—Evergreen forest;
1200–1520 m
elev.
Phenology
:—Flowering in September–October. Fruiting in September–December.
Notes
:—This species is previously considered as a member of genus
Uraria
. It is treated here as a member of
Sohmaea
because it has a straight pod when mature.
Desmodium hispidum
Franch.
is presumably described from a single specimen.
Holotype
of this name is kept at P, whereas its fragments of leaflet and inflorescence along with a photograph of
holotype
itself are kept at A. These fragments should be an isotype in accordance with Art. 8.3, Ex. 8 of ICN (
Turland
et al.
, 2018
).
Specimens examined
:—
THAILAND
.
Chiang Mai
:
Samoeng
,
Yang Moen
,
9 Oct. 1998
,
Maxwell
98-1084
(
BKF
)
,
Doi Chiang Dao Wildlife Sanctuary
,
26 Sept. 1994
,
Nanakorn
et al. 18948
(
QBG
)
,
Chiang Dao
,
Mae Taman
,
27 Sept. 1994
,
Nanakorn
et al. 1932
(
QBG-2
sheets),
Doi Chiang Dao Wildlife Sanctuary
,
20 Dec. 2014
,
Saisorn
343
(
KKU
)
,
Mueang
Chiang Mai
,
Doi Pui village
,
26 Dec. 2014
,
Saisorn
360
(
KKU
)
&
Chiang Dao
,
Tokaew
&
Chantaranothai
854
(
BKF
,
KKU
)
;
Chiang Rai
:
Doi Luang National Park
,
26 Oct. 1997
,
Maxwell
97-1211
(
BKF
)
;
Lampang
:
Wang Nuea
,
Doi Luang National Park
,
7 Nov. 1998
,
Petrmitr
326
(
BKF
)
.