On Venezuelan pholcid spiders (Araneae, Pholcidae)
Author
Huber, Bernhard A.
33607F65-19BF-4DC9-94FD-4BB88CED455F
Zoological Research Museum Alexander Koenig, Bonn, Germany.
b.huber@leibniz-zfmk.de
Author
Villarreal, Osvaldo
679C385E-B068-4351-9D2F-97753E534C26
Museo del Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela. & Museu Nacional / UFRJ, Rio de Janeiro, Brazil.
osvaldovillarreal@gmail.com
text
European Journal of Taxonomy
2020
2020-10-01
718
1
317
journal article
10.5852/ejt.2020.718.1101
4069574
F9E9A91E-488C-4DB1-9361-E788E9AC5BC1
Modisimus repens
Huber
sp. nov.
urn:lsid:zoobank.org:act:
17A58690-0B66-4074-8E5C-3CFDCE23325F
Figs 721–724
,
730–735
,
739–741
, 1023, 1058
Diagnosis
Distinguished from known congeners by morphology of procursus (
Figs 733–734
; widened in midsection, with slender dorsal process and membranous distal fringes), by strong, weakly curved bulbal apophysis (
Figs 730–731
), by armature of male chelicerae (
Fig. 732
; patch of 6–8 stronger hairs on each side on low elevation), and by shape of epigynum and internal female genitalia (
Figs 735
,
739–741
; rectangular epigynum with semicircular whitish area posteriorly; pair of spherical pore plates). From many congeners also by strongly marked femora in males and females (
Figs 721–724
; many ventral dots rather than just subdistal dark rings).
Etymology
The species name (Latin: unexpected) refers to our surprise in finding this ‘regular’ long-legged
Modisimus
in
Venezuela
; adjective.
Type
material
VENEZUELA
–
Falcón
•
♂
holotype
,
ZFMK
(Ar 22062),
Península de Paraguaná
,
Cerro Santa Ana
(
11.8080° N
,
69.9452° W
),
220 m
a.s.l.
,
17 Nov. 2018
(
B.A. Huber
,
O. Villarreal
M.)
.
Other material examined
VENEZUELA
–
Falcón
•
4 ♂♂
,
4 ♀♀
,
ZFMK
(Ar 22063), and
3 ♂♂
,
2 ♀♀
,
1 juv.
in pure ethanol,
ZFMK
(Ven18-189), same collection data as for holotype
.
Description
Male
(
holotype
)
MEASUREMENTS. Total body length 1.9, carapace width 0.85. Distance PME–PME
70 µm
; diameter PME
70 µm
; distance PME–ALE
80 µm
; AME absent. Leg 1: 14.7 (3.9 +0.4+3.7 +5.7+1.0), tibia 2: 2.4, tibia 3: 1.9, tibia 4: 2.3; tibia 1 L/d: 41; all femora approximately same width.
COLOR (in ethanol). Carapace pale ochre-yellow, with three pairs of brown marks and brown median band including ocular area; clypeus dark brown; sternum light brown laterally, whitish medially; legs ochre-yellow, femora with numerous dark ventral marks (in live specimens clearly visible also in dorsal view) and subdistal dark ring, tibiae with proximal and subdistal dark rings; abdomen pale bluish, with many darker bluish and white marks dorsally and laterally, ventrally with light brown mark in gonopore area.
Figs 730–735.
Modisimus repens
Huber
sp. nov.
; from Falcón, Cerro Santa Ana (type locality; ZFMK Ar 22063).
730–731
. Left male pedipalp, prolateral and retrolateral views.
732
. Male chelicerae, frontal view.
733–734
. Left male palpal tarsus and procursus, prolateral and retrolateral views.
735
. Cleared female genitalia, dorsal view. Abbreviations: ba=bulbal apophysis; p=procursus; pp =pore plate; r= receptacle. Scale lines: 0.2 mm.
BODY. Habitus as in
Figs 721–722
. Ocular area moderately raised. Carapace with distinct thoracic groove. Clypeus unmodified. Sternum wider than long (0.58/0.38), unmodified. Abdomen oval.
CHELICERAE. As in
Fig. 732
, with patch of 6–8 stronger hairs on each side on low elevation.
PALPS. As in
Figs 730–731
; coxa with retrolateral apophysis; trochanter barely modified; femur with low retrolateral process proximally and strong ventral apophysis distally; tarsus with conical dorsal process, procursus (
Figs 733–734
) widened in mid-section, with slender dorsal process and distal membranous fringes; genital bulb with strong, weakly curved distal apophysis.
LEGS. Without spines; with curved hairs on tibiae and metatarsi 1–2; with vertical hairs in high density on all femora; retrolateral trichobothrium of tibia 1 at 19%; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~15 pseudosegments, distally distinct.
Male
(variation)
Tibia
1 in
seven males (including
holotype
): 3.6–4.4 (mean 3.9). Some males with curved hairs also on metatarsus 3.
Female
In general similar to male (
Figs 723–724
) but without (or very few) vertical hairs on femora and with lighter sternum (only laterally anteriorly slightly darker brown). Tibia
1 in
six females: 2.4–2.8 (mean 2.6). Epigynum (
Fig. 739
) relatively small, rectangular brown plate with pair of dark marks and semicircular whitish area posteriorly; internal bluish ‘valve’ variably visible in uncleared specimens. Internal genitalia (
Figs 735
,
740–741
) with median receptacle and pair of spherical pore plates, each connected to tapering lateral process.
Figs 736–741.
Modisimus
Simon, 1893
; epigyna, ventral views and cleared female genitalia, ventral and dorsal views.
736–738
.
M. simoni
Huber, 1997
; from Bolívar, Ciudad Guayana (ZFMK Ar 22058).
739–741
.
M. repens
Huber
sp. nov.
; from Falcón, Cerro Santa Ana (type locality; ZFMK Ar 22063).
Distribution
Known from
type
locality only, in
Venezuela
,
Falcón
(Fig. 1058).
Natural history
The spiders were found under rocks near the basis of Cerro Santa Ana, in an area dominated by small trees and bushes (Fig. 1023).