On Venezuelan pholcid spiders (Araneae, Pholcidae)
Author
Huber, Bernhard A.
33607F65-19BF-4DC9-94FD-4BB88CED455F
Zoological Research Museum Alexander Koenig, Bonn, Germany.
b.huber@leibniz-zfmk.de
Author
Villarreal, Osvaldo
679C385E-B068-4351-9D2F-97753E534C26
Museo del Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela. & Museu Nacional / UFRJ, Rio de Janeiro, Brazil.
osvaldovillarreal@gmail.com
text
European Journal of Taxonomy
2020
2020-10-01
718
1
317
journal article
10.5852/ejt.2020.718.1101
4069574
F9E9A91E-488C-4DB1-9361-E788E9AC5BC1
Modisimus
Simon, 1893
Notes
The genus
Modisimus
is species rich in
Mexico
, Central America and the Antilles. Unpublished data suggest that the 79 extant species formally described from this region are just a small fraction of the actual diversity. For example, a recent collecting trip in
Mexico
(
Oct. 2019
) resulted in ~33 species of
Modisimus
, of which ~28 (i.e., 85%) are considered new (B.A. Huber, A. Valdez M., unpubl. data); extensive collections in the Caribbean (mostly
2011–2014
) resulted in ~98 species, of which ~72 (i.e., 73%) are considered new (B.A. Huber, I. Agnarsson, G. Binford, unpubl. data).
By contrast, available data suggest that
Modisimus
is poorly represented in South America. Only three species have previously been reported from mainland South America: the pantropical
M. culicinus
(Simon, 1893)
;
M. simoni
Huber, 1997
from
Venezuela
[including its junior synonym
M. minimus
(
González-Sponga, 2009
)
; see below]; and
M. globosus
Schmidt, 1956
from
Colombia
. A fourth species is newly described below. The faunal turnovers seem to lie somewhere in the
Darien
Gap linking
Panama
and
Colombia
and between the Lesser Antilles (which have a rich unpublished fauna) and
Trinidad
(with no representative of
Modisimus
except
M. culicinus
known to exist in collections).
To some degree, this marked turnover is probably artificial. The pholcid fauna of
Colombia
is very poorly known, and the near-coastal regions of
Venezuela
may also be home to further species. It is unknown whether
Modisimus
has been largely replaced by other taxa in South America or whether it has colonized South America via
Panama
and the Lesser Antilles. The apparent absence of relict species (e.g., in caves and arid regions of
Brazil
) suggests that the second scenario is more likely.