Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae) Author Krikken, J. Author Huijbregts, J. text Zootaxa 2017 4238 3 301 334 journal article 36345 10.11646/zootaxa.4238.3.1 030203e2-e6a1-4b59-bfa0-0689bf817b08 1175-5326 345756 D123950C-DF2B-45EE-9966-813339A83491 Onthophagus manguliensis Boucomont, 1914 ( Figs. 8 , 27 , 53–55 ) Material examined. Syntype male ( MNHN ) from Indonesia , labelled: " Sula Mangoli \ Oct.-Novbr. Doherty", "EX MUSEO \ N .VAN DE POLL", " Onthophagus \ manguliensis \ B. \ COTYPE", "MUSEUM PARIS \ 1938 \ COLL. A. BOUCOMONT". Additional material ( RMNH ) with a similar Doherty collection label ( W. Doherty collected on the Sula Islands around 1890); see also note on type status below. The 35 specimens examined are as follows: CENTRAL SULAWESI : BANGGAI DISTRICT - Batui: Seseba Estate, 80 m , 9.xi.1989 , Krikken & Van der Blom, sw31, grassland, cattle excrement extracted, 16 in RMNH - Matanyo Forest, N of Kayutanyo, 110 m , 2–4.xi.1985 , Krikken & Van der Blom, sw19a, multistratal evergreen forest, river, 2 human excrement traps, 12 in RMNH - Matanyo Forest, N of Kayutanyo, 110 m , 2–4.xi.1989 , Krikken & Van der Blom, sw19b, multistratal evergreen forest, river, 2 fish traps, 2 in RMNH CENTRAL SULAWESI : PALU REGION - Tawaeli-Toboli road (km 29), 250 m , 20–22.xii.1985 , J. Krikken, pw68a, second-growth forest/coffee, near rivulet, 2 human excrement traps, 2 in RMNH SOUTHEAST SULAWESI : KOLAKA DISTRICT - Watuwila Mosquito Camp, 1150 m , 13–15.x.1989 , Krikken & Van der Blom, sw11a, multistratal evergreen forest, 2 human excrement traps, 1 in RMNH SULA ISLANDS - Mangoli, ix-x, Doherty, 1 in RMNH - Mangoli, x-xi, Doherty, 1 in MNHN (syntype) Description. Body length 4–5 mm . Colour symmetrically patterned brown and orange-yellow, slightly variable, largely matt due to microreticulation. Dorsal side with numerous distinct pale-yellow setae. Clypeus anteromedially with pair of apically round denticles, emargination moderately deep; clypeogenal transition continuous at border; low clypeofrontal ridge present, slightly arcuate; paraocular protrusions between anterior border of eyes slight, more distinct in female; clypeus rugulate-punctate, shiny; remainder of head with sparse, coarse seta-bearing punctures and microreticulation ( x60 ). Pronotum slightly, evenly convex; anterior section of lateral border slightly round, posterior section of lateral border scarcely sinuate; punctation double; entire pronotal derm with abundant, seta-bearing, ocellate primary punctation; setae approximately 3 times puncture diameters, punctures mostly separated by 23 diameters; punctures on shallowly depressed, matt basomedian area slightly less pronounced. Elytral striae discally well defined, with distinct punctures, mostly separated by 2–4 times their diameter, slightly crenulated on interstriae; interstriae almost flat, microreticulate ( x40 ), with numerous seta-bearing granules (asperate, x40 ) arranged in indistinct rows; peripheral surfaces of elytra with distinct yellow patches (like humerus, posterior declivity, etc .), remainder brown. Antennal club gray yellowish brown. Metaventrite disc with numerous seta-bearing punctures on anterior lobe; lateral wings with sparse, seta-bearing, subocellate punctures, microreticulate. Abdominal ventrites each with transverse row of seta-bearing punctures. Pygidium yellow, matt, virtually flat, subtriangular; surface with sparse, seta-bearing punctures; base with transverse ridge. Protibia with 3+(1) larger denticles, slightly curved, proximal serration with approximately 10 small denticles; spur elongate-acuminate. Femora largely yellow-brown. Mesotibiae and metatibiae and their spurs unmodified. Metatarsomere 1 and spur almost equal in lenth, shorter than tarsomeres 2–5 combined. Aedeagus, Fig. 55 . Variation, sexual dimorphism, type status. Size slightly variable—Boucomont (1914: 23) mentioned “3 ½– 4 ½ mm”. Colour pattern also only slightly variable. Female similar to male, as shown in Figs. 53–54 . Clearly Boucomont had more specimens before him when he wrote the original description, but lacking details we prefer to maintain their syntype (“cotype”) status. Especially since O . manguliensis -like material requiring study was recently reported from other Moluccan islands (Huijbregts & Krikken 2012).