World Catalogue of the Druid Flies (Diptera: Schizophora: Clusiidae) Author Owen Lonsdale text Zootaxa 2017 2017-10-12 4333 1 1 85 journal article 31851 10.11646/zootaxa.4333.1.1 14597991-8ac8-4290-b2c8-133e4c4329d0 1175-5326 1010066 47AB9D4D-16FF-4A7E-B73F-2A65C2841FF4 Genus HENDELIA Czerny Hendelia Czerny 1903 : 83 . Type species, Hendelia beckeri Czerny 1903 , by monotypy. Gender feminine. Becker 1905 : 39 [catalogue, Palaearctic]; Kertész 1906 : 320 [key]; Williston 1908 : 320 [comparison to Heteroneura ]; Melander & Argo 1924 : 27 [family and genus revision]; Czerny 1928 : 4 [family and genus revision, Palaearctic]; Schulze et al . 1930 : 1517 [nomenclator]; Neave 1939b : 617 [nomenclator]; Hennig 1952 : 227 [discussion of immature stages in Clusiidae ]; Frey 1960 : 11 (as genus and subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]; Trojan 1962 : 21 [key, Poland]; Soós 1963a : 177 [redefinition, key, relationship to Prohendelia and Parahendelia ], 1964a: 7 [key], 1981: 96 [Hungary, key], 1984: 11 [catalogue, Palaearctic]; Cole & Schlinger 1969 : 426 [citation]; Sasakawa 1977 : 238 [catalogue, Orient], 2011b: 13 [Oriental fauna]; Ferrar 1987 : 122 [immature stages, illustrations of larva and puparium]; Stackelberg 1988 : 505 [key, European USSR]; Greve 2005 : 3 [Norway]; Sueyoshi 2006 : 4 [Japan], 2014: 619 [as genus and subgenus, catalogue, Japan]; Papp et al . 2006 : 190 [identified and unidentified species (including possible new subgenus), Thailand]; Lonsdale & Marshall 2007b : 120 [review, biology, redefinition, comparison to Clusiodes , synonymy of Aristohendelia Frey, Burmanochorea Frey , Labomyia Frey , Parahendelia Frey , Prohendelia Frey, Microclusaria Frey, Steleoclusiodes Frey and Xenoclusia Frey , maps for Palaearctic species], 2008b: 5 [Fiji], 2011 [revision, New world], 2010: 1044 [Manual of Central American Diptera ]; Lonsdale 2009b : 230 [key to genera in Australia]; Lonsdale et al. 2010: 267 [family revision, key]; Kahanpää 2014 : 288 [checklist, Finland]. Labomyia Frey 1928a : 14 . Type species, Labomyia mirabilis Frey 1928 , by original designation. Gender feminine. Neave 1939b : 842 [nomenclator]; Hennig 1952 : 227 [discussion of immature stages in Clusiidae ]; Frey 1960 : 26 [catalogue, world]; McAlpine 1960 : 67 [key]; Soós 1964a : 6 [key], 1968: 4 [catalogue, Americas south of USA ]; Caloren & Marshall 1998 : 263 [as junior synonym of Clusiodes ]; Lonsdale & Marshall 2007b : 120 [as synonym of Hendelia ]. Aristohendelia Frey 1960 : 14 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, World]. Gender feminine. Type species, Hendelia extensicornis Frey 1960 , by original designation. Sasakawa 1977 : 238 [catalogue, Orient]; Lonsdale & Marshall 2007b : 120 [as synonym of Hendelia ]. Burmanochorea Frey 1960 : 14 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Type species, Hendelia variabilis Frey 1960 , by original designation. Gender feminine. Sasakawa 1977 : 238 [catalogue, Orient]; Lonsdale & Marshall 2007b : 120 [as synonym of Hendelia ]. Prohendelia Frey 1960 : 13 (as subgenus) [family and genus revision, key to subgenera and species, family and genus revision, Indoaustralia, catalogue, world]. Type species, Hendelia orientalis Frey 1960 , by original designation. Gender feminine. Soós 1963a : 177 [new status as genus, key, relationship to Hendelia and Parahendelia ], 1964a: 7 [key]; Sasakawa 1977 : 238 [catalogue, Orient]; Balthazar 1990 : 417 [catalogue, Philippines ]; Lonsdale & Marshall 2007b : 120 [as synonym of Hendelia ]. Microclusiaria Frey 1960 : 19 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Gender feminine. Type species, Clusiodes obscuripennis Frey 1960 , by original designation. Sasakawa 1977 : 237 [catalogue, Orient]; Balthazar 1990 : 417 [catalogue, Philippines ]; Papp et al . 2006 : 190 [identified and unidentified species; Thailand ]. Lonsdale & Marshall 2007b : 120 [as synonym of Hendelia ]. Steleoclusiodes Frey 1960 : 18 (as subgenus) [family and genus revision, key to subgenera and species, Indoaustralia, catalogue, world]. Type species, Clusiodes nigrifrons Frey 1960 , by original designation. Gender masculine. Sasakawa 1977 : 237 [catalogue, Orient], 1987: 101 [key, Orient/Pacific]; Lonsdale & Marshall 2007b : 120 [as synonym of Hendelia ]. Xenoclusia Frey 1960 : 21 [catalogue, world]. Type species: Xenoclusia latifrons Frey 1960 [by original designation]. Gender feminine. Sasakawa 1977 : 239 [catalogue, Orient]; Soós 1964a : 7 [key]; Balthazar 1990 : 417 [catalogue, Philippines ]; Lonsdale & Marshall 2007b : 120 [as synonym of Hendelia ]. Parahendelia McAlpine 1960 : 83 . Type species, Parahendelia latifrons McAlpine 1960 , by original designation (= Hendelia armiger Lonsdale & Marshall ). Gender feminine. Soós 1963a : 179 [key, relationship to Hendelia and Prohendelia ], 1964a: 4 [key]; Pitkin & Evenhuis 1989 : 536 [catalogue, Australasia/Pacific]. Marshall 2000 : 5 [as junior synonym of Clusiodes ); Lonsdale & Marshall 2007b : 120 [as synonym of Hendelia ]. Prophedelia . Misspelling of Prohendelia . Hua 2006 : 160 [catalogue—not known from China ]. Lonsdale & Marshall 2007b [citation]. aberrans (Frey) . OR: Malaysia ( Sabah ) , Philippines. Clusiodes aberrans Frey 1928b : 106 . Philippines. Oriental Mindoro Province: “Calopan” [= Calapan]. HT ♀, depository unknown [originally deposited UZMH]. Delfinado & Hardy 1971 : 29 [catalogue Philippine types]. Clusiodes ( Microclusiaria ) aberrans . Frey 1960 : 19 [Philippines, catalogue, world]; Sasakawa 1977 : 237 [catalogue, Orient]; Balthazar 1990 : 417 [catalogue, Philippines]. Clusiodes ( Clusiaria ) aberrans . Sasakawa 1987 : 103 [key, Orient/Pacific]. Hendelia aberrans . Lonsdale & Marshall 2007b : 147 [generic combination]; Sasakawa 2011b : 14 [ Malaysia ]. aenigma Frey. OR: Burma. Hendelia ( Burmanochorea ) aenigma Frey 1960 : 15 . Burma. NE Burma: Kambaiti, 2000m . HT ♀ UZMH. Frey 1960 : 15 [catalogue, world]; Sasakawa 1977 : 238 [catalogue, Orient]. Hendelia aenigma . Lonsdale & Marshall 2007b : 147 [list]. amerinx Lonsdale & Marshall. AU: Fiji. Hendelia amerinx Lonsdale & Marshall 2008b : 7 . Fiji. Viti Levu: Naitasiri Province , 4km WSW Colo-i-Suva Village , Mt. Nakobalevu , 372 m . HT ♂ FNIC . Lonsdale & Marshall 2008b : 7 [illustrations of male genitalia]. angulosa (Sueyoshi) . PA: Japan. Clusiodes angulosus Sueyoshi 2006 : 6 . Japan. Honshu: Sarugajo, Kitaibaraki City, Ibaraki Prefecture, Honshu. HT ♂ BLKU. Sueyoshi 2006 : 6 [map, illustrations of head and male and female genitalia, photo of wing], 2011: 7 [Japan, pictoral key], 2014: 618 [catalogue, Japan]; Lonsdale & Marshall 2007b : 154 [ lapsus ]. Hendelia angulosa Lonsdale & Marshall 2007b : 147 [generic combination]; Sueyoshi 2012 : 27 [Japan]. arguta (McAlpine) . AU: Australia (New South Wales, Victoria). Clusiodes arguta McAlpine 1960 : 82 . Australia. New South Wales: Blue Mountains, Wentworth Falls. HT ♂ AMS. Clusiodes ( Clusiodes ) arguta . Sasakawa 1987 : 102 [key, Orient/Pacific]. Clusiodes argutus . Pitkin & Evenhuis 1989 : 535 [catalogue, Australasia/Pacific]. Hendelia arguta . Lonsdale & Marshall 2007b : 147 [generic combination]. armiger Lonsdale & Marshall. AU: Australia (New South Wales, Queensland). Parahendelia latifrons McAlpine 1960 : 84 . Australia. New South Wales: Blue Mountains, Wentworth Falls. HT ♂ AMS. McAlpine 1960 : 84 [illustrations of head], 1976: 75 [discussion of vibrissal structure]; Pitkin & Evenhuis 1989 : 536 [catalogue, Australasia/Pacific]. Clusiodes latifrons . Marshall 2000 : 3 [generic combination, male agonistic behaviour, photos]. Hendelia armiger Lonsdale & Marshall 2007b : 118 [generic combination, replacement for name preoccupied by Frey (1960) ]. beckeri Czerny. PA: Austraia and central Europe to Norway , Finland , eastern and northern coasts of the Black Sea, central and eastern Russia , Japan , China ( Sichuan , Zeijiang)*, South Korea *. Hendelia Beckeri Czerny 1903 : 84 . Austria. Bad Hall. ST ♂♀, depository unknown. Czerny 1903 : 84 [illustrations of body and heads of both sexes], 1928: 4 [redescription, illustrations of head and body, redescription of females as nigripalpis Czerny ]; Becker 1905 : 39 [catalogue, Palaearctic]. Hendelia beckeri . Melander & Argo 1924 : 27 [summary of original description]; Tuomikoski 1936 : 183 [ Finland ]; Trojan 1962 : 21 [key, redescription, illustrations of male and female heads]; Sasakawa 1963: 18 [ Japan ]; Hackman 1980 : 148 [checklist, Finland ]; Soós 1981 : 97 [ Hungary , illustrations of habitus, head], 1984: 11 [catalogue, Palaearctic]; Greve 1983 : 55 [habitat, Norway ], 2005: 3 [considered rare, Norway ]; Greve & Midtgaard 1986 : 91 [ Norway ]; Zuska 1987 : 235 [ Czech Republic ]; Mamaev 1987 : 115 [Russian Far East]; Stackelberg 1988 : 505 [key, illustration of female head, European USSR]; Ceianu 1989 : 29 [catalogue, Romania ]; Roháček 1995 : 125 [puparia in decaying alder and birch, adults on aspen trunks, biology and habitat, Czech and Slovak Republics], 1997: 75 [checklist, Czech and Slovak republics], 2009: 219 [photo of male, Slovakia ], 2012: 99 [central Slovakia ], 2013: 128 [central Slovakia ]; Merz 1998 : 276 [checklist, Switzerland ]; Yagi & Sawada 2000: 40 [Mamaev collection holdings]; Gärdenfors 2000 : 222 [Red List of Swedish species]; Roháček et al . 2005: 280 [catalogue, habitat, considered local and rare in Czech Republic , Czech Republic ]; Barták & Kubík 2005: 280 [ Czech Republic ]; Sueyoshi 2006 : 14 [ Japan ], 2011: 6 [ Japan , pictoral key], 2012: 28 [ Japan ], 2014: 619 [catalogue, Japan ]; Lonsdale & Marshall 2007b : 147 [biology, illustrations of genitalia and heads, relationships]; Schacht et al. 2007 : 66 [ Germany ]; Nilsson 2009 : 5 [ Sweden , illustration of female]; Adaschkiewitz 2009 : 287 [ Germany ]; Kahanpää 2014 : 288 [checklist, Finland ]. Hendelia nigripalpis Czerny 1928 : 5 [illustration of head]. Austria. ST ♀ [= ♀ ST of H. Beckeri Czerny ], depository unknown. Frey 1928c : 108 [catalogue, Finland]; Tuomikoski 1936 : 183 [as synonym of beckeri Czerny ]. Hendelia (Hendelia) beckeri . Frey 1960 : 13 [redescription, catalogue, World]. bisetosa (Mamaev) . PA: Russia (Far East). Clusiodes bisetosa Mamaev 1974 : 115 . Russia . Far East: Kedrovaja Pad Nature Reserve. HT ♂ MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1974 : 115 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]. Hendelia bisetosa . Lonsdale & Marshall 2007b : 147 [generic combination, map]. bismarckensis (Sasakawa) . AU: Papua New Guinea ( Lavongai Island ). Clusiodes bismarckensis Sasakawa 1974 : 160 . Papua New Guinea. Bismarck Archipelago: Lavongai Island, Banatam. HT ♂ UZMC. Sasakawa 1974 : 160 [illustrations of male genitalia], 1987: 101 [key, Orient/Pacific]; Pitkin & Evenhuis 1989 : 535 [catalogue, Australasia/Pacific]. Hendelia bismarckensis . Lonsdale & Marshall 2007b : 147 [generic combination]. boliviensis Lonsdale & Marshall. NT: Bolivia. Hendelia boliviensis Lonsdale & Marshall 2011: 4 . Bolivia . La Paz Department : Caranavi, ca. 10km NW road to ENTEL tower, 1400 m , 15°46′35ʺS, 67°35′48ʺW. HT ♀ ANCB. Lonsdale & Marshall 2011: 4 [illustration of female genitalia]. campa Lonsdale & Marshall. NT: Mexico. Hendelia campa Lonsdale & Marshall 2011: 4 . Mexico. Veracruz State: Xalapa. HT ♂ TAUI . Lonsdale & Marshall 2011: 4 [illustrations of male genitalia, photo]. chaetostylotis (Sasakawa) . AU: Indonesia ( West Papua ). Clusiodes ( Clusiodes ) chaetostylotis Sasakawa 1987 : 103 . Indonesia . West Papua : Kebar Valley, Vogelkop, W of Manokwari, 550 m . HT ♂ BPBM [ Type No. 13580]. Sasakawa 1987 : 103 [illustration of male genitalia]. Hendelia chaetostylotis . Lonsdale & Marshall 2007b : 147 [generic combination]. clara (McAlpine) . AU: Australia ( Queensland ). Clusiodes clara McAlpine 1960 : 83 . Australia. Queensland: Silkwood, near Innisfail. HT ♀ AMS. Clusiodes ( Clusiodes ) clara . Sasakawa 1987 : 103 [key, Orient/Pacific]. Clusiodes clarus . Pitkin & Evenhuis 1989 : 535 [catalogue, Australasia/Pacific]. Hendelia clara . Lonsdale & Marshall 2007b : 147 [generic combination]. dasyta (Sasakawa) . AU: Solomon Islands (New Georgia Island). Clusiodes ( Columbiella ) dasytus Sasakawa 1987 : 113 . Solomon Islands . New Georgia Island: Munda, 1– 30m . HT ♂ BPBM [ Type No. 13587]. Sasakawa 1987 : 113 [illustration of male genitalia]. Hendelia dasyta . Lonsdale & Marshall 2007b : 147 [generic combination]. extensicornis Frey. OR: Burma , Thailand . Hendelia ( Aristohendelia ) extensicornis Frey 1960 : 14 . Burma. NE Burma, Kambaiti, 2000 m . ST 6♂ 3♀ UZMH. Frey 1960 : 14 [illustrations of head, catalogue, world]; Papp et al . 2006 : 191 [Thailand]. Hendelia extensicornis . Lonsdale & Marshall 2007b : 147 [relationships]. femorata (Sasakawa) . AU: Indonesia ( Papua ). Clusiodes ( Clusiodes ) femoratus Sasakawa 1987 : 105 . Indonesia. Papua Province : Ifar , east end of Cyclops Mts. , 150 m . HT ♀ BPBM [Type No. 13581]. Hendelia femorata . Lonsdale & Marshall 2007b : 147 [generic combination]. formosana (Hennig) . OR: Taiwan. Clusiodes ( Microclusiaria ) formosana Hennig 1938 : 128 . Taiwan. Paiwan District: northern Paroe. HT ♂ DEI. Clusiodes formosana . Hennig 1941 : 146 [Taiwan]; Hua 2006 : 160 [catalogue—not known from China]. Clusiodes ( Microclusiaria ) formosana . Frey 1960 : 19 [catalogue, world]; Sasakawa 1977 : 237 [catalogue, Orient]. Clusiodes ( Clusiaria ) formosana . Sasakawa 1987 : 103 [key, Orient/Pacific]. Hendelia formosana . Lonsdale & Marshall 2007b : 147 [generic combination]. freyi (Soós) . OR: Taiwan. Prohendelia ( Prohendelia ) freyi Soós 1963a : 179 . Taiwan. Kosempo. HT ♂ HNHM. Soós 1963a : 179 [illustrations of head]; Sasakawa 1977 : 239 [catalogue, Orient]. Hendelia freyi . Lonsdale & Marshall 2007b : 147 [generic combination]. fulvifasciata Sasakawa. OR: Malaysia ( Sabah ). Hendelia fulvifasciata Sasakawa 2011b : 14 . Malaysia. Sabah: 19 km N of Kalabakan, forest camp. HT ♂, BPBM [Type No. BPBM 17352]. Sasakawa 2011b : 14 [illustrations of male genitalia]. geniculata Sasakawa. OR: Malaysia ( Sabah ). Hendelia geniculata Sasakawa 2011b : 16 . Malaysia. Sabah: 9.8 km SW of Tenom, forest camp. HT ♂, BPBM [Type No. BPBM 17355]. Sasakawa 2011b : 16 [illustrations of male genitalia]. gladiator (McAlpine) . AU: Australia (New South Wales, Queensland, Tasmania, Victoria). Clusiodes gladiator McAlpine 1960 : 81 . Australia. New South Wales: National Park, south of Sydney. HT ♂ AMS. McAlpine 1960 : 81 [illustrations of head and wing], 1976: 75 [discussion of spiral vibrissae and sexual dimorphism, illustrations of head and male postabdomen]; Pitkin & Evenhuis 1989 : 535 [catalogue, Australasia/Pacific]; Marshall 2000 : 3 [male agonistic behaviour, photos]. Clusiodes ( Clusiodes ) gladiator . Sasakawa 1987 : 102 [key, Orient/Pacific]. Hendelia gladiator . Lonsdale & Marshall 2007b : 147 [generic combination]; Lonsdale et al. 2010: 241 [genes sequenced for phylogenetic analysis]. heliconiae Lonsdale & Marshall. NT: Bolivia , Costa Rica , Ecuador . Hendelia heliconiae Lonsdale & Marshall 2011: 6 . Ecuador. Pichincha Province: Maquipucuna Biological Research Station, river trail, 1200 m . HT ♂ QZAC. Lonsdale & Marshall 2010 : 1046 [illustration of female genitalia as Hendelia sp.], 2011: 6 [illustrations of male and female genitalia, photo]. iotoides (Sasakawa) . AU: Malaysia ( Sabah ). Clusiodes iotoides Sasakawa in Sasakawa & Pong 1990 : 55 . Malaysia. Sabah: Sepilok Forest Reserve. HT ♂ FRC. Sasakawa & Pong 1990 : 55 [illustrations of male genitalia]. Hendelia iotoides . Lonsdale & Marshall 2007b : 147 [generic combination]. kinetrolicros (Caloren & Marshall) . NT: Costa Rica , Mexico ( Chiapas , Veracruz ). Clusiodes kinetrolicros Caloren & Marshall 1998 : 295 . Costa Rica. Guanacaste Province : Guanacaste Conservation Area, Cacao Field Station , 900 m . HT ♂ CNC . Caloren & Marshall 1998 : 295 [illustrations of head, wing and male and female genitalia]. Hendelia kinetrolicros . Lonsdale & Marshall 2007b : 147 [generic combination], 2010: 1043 [Manual of Central American Diptera ], 2011: 6 [photo, reproduction of genitalic figures]. laosica Sasakawa. OR: Laos. Hendelia laosica Sasakawa 2011b : 16 . Laos. Vientian: Gi Sion Village, de Tha Ngone. HT ♂, BPBM [Type No. BPBM 17353]. Sasakawa 2011b : 16 [illustrations of male genitalia]. lateralis (Sasakawa) . OR: Philippines. Prohendelia lateralis Sasakawa 1974 : 153 . Philippines . Palawan : Pinigisan, 600 m . HT ♂ UZMC. Sasakawa 1974 : 153 [illustrations of head and male genitalia], 1977: 239 [catalogue, Orient]. Hendelia (Prohendelia) lateralis . Balthazar 1990 : 417 [catalogue, Philippines ]. Hendelia lateralis . Lonsdale & Marshall 2007b : 147 [genus redefinition]. latifrons (Frey) . OR: Philippines. Xenoclusia latifrons Frey 1960 : 21 . Philippines. Mindanao: Suriago. HT ♀ UZMH. Frey 1960 : 21 [catalogue, world]; Delfinado & Hardy 1971 : 29 [catalogue Philippine types]; Balthazar 1990 : 417 [catalogue, Philippines]. Hendelia latifrons . Lonsdale & Marshall 2007b : 147 [generic combination, illustration of head]. marginalis (Sasakawa) . AU: Papua New Guinea (Bougainville Island), Indonesia (Biak Island). Clusiodes ( Clusiodes ) marginalis Sasakawa 1987 : 106 . Papua New Guinea. Bougainville Island: Kokure, 690 m . HT ♂ BPBM [Type No. 13582]. Sasakawa 1987 : 106 [illustration of male genitalia]. Hendelia marginalis . Lonsdale & Marshall 2007b : 147 [generic combination]. masneri Lonsdale & Marshall. NT: Costa Rica. Hendelia masneri Lonsdale & Marshall 2011: 8 . Costa Rica. Puntarenas Province : Monteverde Research Station , 1550 m . HT ♂ INBio . Lonsdale & Marshall 2011: 8 [illustrations of male genitalia]. megaspilos (McAlpine) . AU: Australia (New South Wales, Queensland). Clusiodes megaspilos McAlpine 1960 : 83 . Australia. New South Wales : Blue Mountains , Wentworth Falls . HT ♂ AMS. Sasakawa 1987 : 102 [key, Orient/Pacific]. Hendelia megaspilos . Lonsdale & Marshall 2007b : 147 [generic combination]. megastylotis (Sasakawa) . AU: Indonesia ( West Papua ) , Papua New Guinea. Clusiodes ( Clusiodes ) megastylotis Sasakawa 1987 : 107 . Indonesia . West Papua : Manokwari, Vogelkop, 75 m . HT ♂ BPBM [ Type No. 13583]. Sasakawa 1987 : 107 [illustrations of male genitalia]. Hendelia megastylotis . Lonsdale & Marshall 2007b : 147 [generic combination]. melanospilus (Sasakawa) . AU: Papua New Guinea ( Bismarck Isl. , Morobe ) . Clusiodes ( Clusiodes ) melanospilus Sasakawa 1987 : 109 . Papua New Guinea. NE New Guinea: Morobe District, Wau, 1200 m . HT ♂ BPBM [Type No. 13584]. Pitkin & Evenhuis 1989 : 535 [catalogue, Australasia/Pacific]. Hendelia melanospilos . Misspelling of melanospilus . Lonsdale & Marshall 2007b : 147 [generic combination]. mirabilis (Frey) . NT: Bolivia , Costa Rica , Ecuador , Peru . Labomyia mirabilis Frey 1928a : 16 . Peru. Eastern Andes, Roque. LT 1♂ UZMH [accidental designation of lectotype by assumption of holotype (IZCN (1999) Art. 74.6) by Caloren & Marshall (1998) ; 2♂ ST originally described]. Frey 1960 : 26 [illustration of head, catalogue, world] Clusiodes mirabilis . Caloren & Marshall 1998 : 297 [generic combination, redescription, new record ( Peru ), illustrations of head, wing and male and female genitalia]. Hendelia mirabilis . Lonsdale & Marshall 2007b : 147 [generic combination, habitus photos], 2010: 1043 [Manual of Central American Diptera ], 2011: 9 [description of variation, photos, illustrations of male genitalia, new records ( Bolivia , Costa Rica , Ecuador )]. napo (Caloren & Marshall) . NT: Ecuador , Venezuela . Clusiodes napo Caloren & Marshall 1998 : 305 . Ecuador. Napo Province: Baeza, 2000 m . HT ♂ DEBU. Caloren & Marshall 1998 : 305 [illustrations of wing and male and female genitalia]. Hendelia napo . Lonsdale & Marshall 2007b : 147 [generic combination], 2011: 12 [reproduction of genitalic illustrations, new records ( Venezuela )]. nigriceps (McAlpine) . AU: Australia ( New South Wales ). Parahendelia nigriceps McAlpine 1960 : 85 . Australia. New South Wales: Blue Mountains, Mt. Wilson. HT ♂ AMS. McAlpine 1960 : 85 [illustration of head], 1976: 75 [discussion of spiral vibrissae]; Pitkin & Evenhuis 1989 : 536 [catalogue, Australasia/Pacific]. Clusiodes nigriceps . Marshall 2000 : 3 [generic combination, male agonistic behaviour, photos]. Hendelia nigriceps . Lonsdale & Marshall 2007b : 147 [generic combination]. nigrifrons (Frey) . OR: Burma. Clusiodes ( Steleoclusiodes ) nigrifrons Frey 1960 : 18 . Burma. NE Burma: Kambaiti, 2000 m . ST 7♂ 3♀ UZMH. Frey 1960 : 18 [catalogue, world]; Sasakawa 1977 : 237 [catalogue, Orient]; Papp et al . 2006 : 190 [Thailand]. Hendelia nigrifrons . Lonsdale & Marshall 2007b : 147 [generic combination]. nigripalpus Lonsdale & Marshall. NT: Bolivia. Hendelia nigripalpus Lonsdale & Marshall 2011: 12 . Bolivia . La Paz Department : Caranavi, ca. 10 km NW, road to ENTEL tower, 1400 m , 15°46′35ʺS, 67°35′48ʺW. HT ♂ ANCB. Lonsdale & Marshall 2011: 12 [illustrations of male genitalia]. notata (Sasakawa) . OR: Malaysia ( Sarawak ). Clusiodes ( Clusiodes ) notatus Sasakawa 1987 : 111 . Malaysia . Sarawak : Kampong , Sadong , Tapuh , 300–450 m . HT ♂ [ Type No. 13585], depository unknown [originally deposited in BPBM ] . Hendelia notata . Lonsdale & Marshall 2007b : 147 [generic combination]; Sasakawa 2011b : 18 [ Malaysia , illustration of male genitalia]. obscuripennis (Frey) . OR: Burma , Thailand . Clusiodes ( Microclusiaria ) obscuripennis Frey 1960 : 19 . Burma . Tenasserim, Sukli, 75 km E of Moulmein, 600 m . HT ♀ , depository unknown [originally deposited at UZMH]. Frey 1960 : 19 [catalogue, world]; Sasakawa 1977 : 237 [catalogue, Orient]; Papp et al . 2006 : 190 [ Thailand ]. Clusiodes ( Clusiaria ) obscuripennis . Sasakawa 1987 : 103 [key, Orient/Pacific]. Hendelia obscuripennis . Lonsdale & Marshall 2007b : 147 [generic combination]. orientalis (Frey) . OR: Burma. Hendelia ( Prohendelia ) orientalis Frey 1960 : 14 . Burma. NE Burma : Kambaiti , 2000 m . ST 1♂ 2♀ UZMH . Frey 1960 : 14 [catalogue, world]. Prohendelia orientalis . Soós 1963a : 182 [key, illustrations of head]; Sasakawa 1977 : 239 [catalogue, Orient]. Hendelia orientalis . Lonsdale & Marshall 2007b : 147 [generic combination]. orientalis form conjuncta Frey. OR : Burma . Hendelia ( Prohendelia ) orientalis form conjuncta Frey 1960 : 14 . Burma . Kambaiti. ST 1♂ 2♀ UZMH . orientalis form obscurior Frey. OR: Burma. Hendelia ( Prohendelia ) orientalis form obscurior Frey 1960 : 14 . Burma. Kambaiti. ST 1♀ UZMH. palliceps Frey. OR: Burma. Hendelia ( Burmanochorea ) palliceps Frey 1960 : 14 . Burma. NE Burma: Kambaiti, 2000 m . ST 5♂ 2♀ UZMH. Frey 1960 : 14 [catalogue, world]; Sasakawa 1977 : 238 [catalogue, Orient]. Hendelia palliceps . Lonsdale & Marshall 2007b : 147 [list]. personata Frey. OR: Burma. Hendelia ( Burmanochorea ) personata Frey 1960 : 15 . Burma. NE Burma: Kambaiti, 2000 m . HT ♂ UZMH. Frey 1960 : 15 [catalogue, world]; Sasakawa 1977 : 238 [catalogue, Orient]. Hendelia personata . Lonsdale & Marshall 2007b : 147 [list]. phrenzinus (Caloren & Marshall) . NT: Costa Rica , Venezuela . Clusiodes phrenzinus Caloren & Marshall 1998 : 307 . Venezuela. Aragua State: Parque Nacional Henry Pittier, Highway Maracay-Choroni, km 19, 1330 m. HT ♂ DEBU. Caloren & Marshall 1998 : 307 [illustrations of wing and male and female genitalia]. Hendelia phrenzinae . Lonsdale & Marshall 2007b : 147 [generic combination, unjustified emendation]. Hendelia phrenzinus . Lonsdale & Marshall 2011: 13 [reproduction of genitalic figures, new record ( Costa Rica )]. philippinensis Sasakawa. OR: Philippines. Hendelia philippinensis Sasakawa 2011b : 18 . Philippines . Luzon: Nueva Vizcaya , Dalton Pass, 915 m . HT ♂ , BPBM [ Type No. BPBM 17350]. Sasakawa 2011b : 18 [illustrations of male genitalia]. plumipes (Sasakawa) . OR: Japan (Ryukyus), Malaysia ( Sabah ) . Clusiodes (Clusiodes) plumipes Sasakawa 1987 : 112 . Malaysia. Sabah: Ranau, 500 m . HT ♀ BPBM [Type No. 13586]. Hendelia plumipes . Lonsdale & Marshall 2007b : 147 [generic combination]; Sasakawa 2011b : 20 [Oriental revision]; Sueyoshi 2014 : 619 [catalogue, Japan]. Clusiodes usikumuri Sueyoshi 2006 : 11 . Japan. Ryuku: Irimote Island, Ushikumori, Taketomi Town. HT ♀ BLKU. Sueyoshi 2006 : 11 [illustrations of head and female genitalia, photo of wing]; Sasakawa 2011b : 20 [as synonym of plumipes ]. Hendelia usikumuri . Lonsdale & Marshall 2007b : 147 [generic combination], 2011: 7 [ Japan , pictoral key]. plumosa (Sasakawa) . OR: Japan (Ryukyus). PA: Japan (widespread). Clusiodes (Clusiodes) plumosus Sasakawa 1964 : 14 . Japan. Honshu: Ashiu, Kyoto. HT ♀ OMNH. Sasakawa 1964 : 14 [lateral illustration], 1990: 540 [description of male]; Soós 1984 : 12 [catalogue, Palaearctic]; Matsumoto & Sasakawa 2006 : 19 [notes holotype deposition in OKAKA]. Clusiodes plumosus . Sueyoshi 2006 : 9 [illustrations of head and male genitalia, photo of wing]. Hendelia plumosa . Lonsdale & Marshall 2007b : 147 [generic combination, map]; Sueyoshi et al. 2009 : 101 [log emergence trap, Japan ]; Sueyoshi 2011 : 8 [ Japan , pictoral key], 2012: 28 [ Japan ], 2014: 619 [catalogue, Japan ]. pterygion (Caloren & Marshall) . NT: Southern Brazil . Clusiodes pterygion Caloren & Marshall 1998 : 299 . Brazil. Rio de Janeiro : Rio de Janeiro . HT ♂ AMNH . Caloren & Marshall 1998 : 299 [illustrations of male genitalia and wing]. Hendelia pterygion . Lonsdale & Marshall 2007b : 147 [generic combination], 2011: 13 [reproduction of genitalic figures]. punctifrons (Frey) . OR: Burma. Clusiodes ( Clusiodes ) punctifrons Frey 1960 : 18 . Burma. NE Burma: Kambaiti, 2000 m . ST 8♂ 7♀ UZMH. Frey 1960 : 18 [catalogue, world]; Sasakawa 1977 : 237 [catalogue, Orient], 1987: 102 [key, Orient/Pacific]. Hendelia punctifrons . Lonsdale & Marshall 2007b : 147 [generic combination]. putealis Lonsdale & Marshall. NT: Costa Rica. Hendelia putealis Lonsdale & Marshall 2011: 15 . Costa Rica. Cartago Province : Cerro de la Muerte La Cangreja , 1950 m . HT ♀ INBio . Lonsdale & Marshall 2011: 15 [illustration of female genitalia, photo]. quatuorsetosa (Mamaev) . PA: Russia (Far East). Clusiodes quatuorsetosa Mamaev 1974 : 114 . Russia . Far East: Ussury Krai. HT ♂ [with puparium] MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1974 : 114 [illustration of surstylus]; Yagi & Sawada 2000: 41 [Mamaev collection holdings]. Hendelia quatuorsetosa . Lonsdale & Marshall 2007b : 147 [generic combination, map]. salebrosa Lonsdale & Marshall. NT: Bolivia. Hendelia salebrosa Lonsdale & Marshall 2011: 15 . Bolivia. La Paz Department : Zongo , 16°06′32ʺS, 68°04′30ʺW. HT ♂ ANCB . Lonsdale & Marshall 2011: 15 [illustrations of male and female genitalia]. saopaulo (Caloren & Marshall) . NT: Brazil ( São Paulo , Rio de Janeiro ). Clusiodes saopaulo Caloren & Marshall 1998 : 303 . Brazil. São Paulo : São Paulo , Jaragua. HT ♂ MNRJ . Caloren & Marshall 1998 : 303 [illustrations of wing and male and female genitalia]. Hendelia saopaulo . Lonsdale & Marshall 2007b : 147 [generic combination], 2011: 16 [reproduction of genitalic figures]. similis Lonsdale & Marshall. AU: Fiji. Hendelia similis Lonsdale & Marshall 2008b : 8 . Fiji . Viti Levu: Nukura Forest, 60– 130m . HT ♂ BPBM [ Type No. 16838]. Lonsdale & Marshall 2008b : 8 [illustrations of egg and male and female genitalia, photo]. spilota Sasakawa. OR: Malaysia ( Sabah ). Hendelia spilota Sasakawa 2011b : 20 . Malaysia. Sabah: 19 km N of Kalabakan. HT ♀, BPBM [Type No. BPBM 17351]. Sasakawa 2011b : 20 [illustrations of male genitalia]. stimulator (Caloren & Marshall) . NE: Mexico ( Tamaulipas ). Clusiodes stimulator Caloren & Marshall 1998 : 301 . Mexico. Tamaulipas State: “ Mpla ”, Gomez Farias , Rancho de Cielo. HT ♂ TAMU . Caloren & Marshall 1998 : 301 [illustrations of male genitalia and wing]. Hendelia stimulator . Lonsdale & Marshall 2007b : 147 [generic combination], 2011: 16 [reproduction of genitalic figures]. unica Mamaev. PA: Russia (Far East). Hendelia ( Burmanochorea ) unica Mamaev 1974 : 112 . Primorsky Krai : Kedroveja [= Kedrovaya] Pad Nature Reserve. HT ♂ MNHA [not in ZIN, as stated in original publication, examined by author]. Mamaev 1987 : 115 [Russian Far East]. Hendelia unica . Lonsdale & Marshall 2007b : 147 [list, map]. variabilis Frey. OR: Burma. Hendelia ( Burmanochorea ) variabilis Frey 1960 : 15 . Burma. NE Burma: Kambaiti, 2000 m . ST 1♂ 3♀ UZMH. Frey 1960 : 15 [catalogue, world]; Sasakawa 1977 : 238 [catalogue, Orient]. Hendelia variabilis . Lonsdale & Marshall 2007b : 147 [list].