The inclusion of Akeassia in Grangea (Asteraceae) and description of a new species from Gabon: Grangea ogoouensis
Author
Beentje, Henk
Author
Lachenaud, Olivier
text
Candollea
2020
2020-11-10
75
2
311
319
journal article
20541
10.15553/c2020v752a12
479dd8b7-51b3-425e-b74b-6ba4a5e46f35
2235-3658
5724909
Key to the species of
Grangea
(Species occurring in
Gabon
are in bold).
1. Pappus a coroniform ring with dentate or laciniate margin (the ring sometimes very short in
G. ceruanoides
but then outer florets in a single series) ....................................... 2
1a. Pappus of free bristles, or bristle-scales briefly connate at base, or absent; outer florets in several series ................. 6
2. Leaves shallowly dentate (teeth to
2 mm
deep), long-attenuate at base; achenes glandular, without hairs; endemic to
Burundi
.......................................................
G. jeffreyana
2a. Leaves pinnatifid, or if shortly toothed then auriculate at base; achenes mostly hairy (sometimes glabrous in
G. maderaspatana
) and often glandular as well .............. 3
3. Capitula 3 – 6 together in terminal corymbs; hairs on achenes simple and uncinate at apex; pappus margin very shortly and irregularly toothed; erect herb;
Angola
,
Zambia
,
Zimbabwe
,
Namibia
,
Botswana
................
G. anthemoides
3a. Capitula, at least the majority of them, solitary; hairs on achenes (when present) either bifid, or if simple then not uncinate; pappus margin laciniate with ± linear teeth; erect or decumbent herbs ....................................................... 4
4. Outer florets in one series; achenes eglandular, compressed; Sahelian region from
Senegal
to
South Sudan
................. ...................................................................
G. ceruanoides
4a. Outer florets in several series; achenes glandular, compressed or quadrangular ................................................ 5
5. Achenes strongly quadrangular in cross section; pappus ring less than
0.5 mm
high; phyllaries
2–3 mm
long; plant low and very compact, c.
10 cm
tall; endemic to
Zambia
......... ...................................................................
G. zambesiaca
5a. Achenes compressed, elliptic and weakly 2(– 4)angled in cross-section; pappus ring to
1.3 mm
high; phyllaries
3–7 mm
long; plant very variable in habit, usually taller; widespread in tropical and subtropical Africa,
Madagascar
, and tropical Asia ................................
G. maderaspatana
6. Marginal florets almost urceolate, often red; phyllaries to
4 mm
; achenes without hairs; pappus absent or consisting of 1–2 free bristles;
Madagascar
........................
G. lyrata
6a. Marginal florets with narrow cylindric base, always yellow; phyllaries to
3 mm
long; achenes with hairs; pappus present (sometimes caducous in
G. gossypina
) ............................ 7
7. Pappus of relatively broad bristle-scales to
1 mm
long, shortly connate at base; leaves shallowly dentate;
Madagascar
...............................................
G. madagascariensis
7a. Pappus of smaller and filiform free bristles; leaves shallowly dentate to pinnatifid ...................................................... 8
8. Capitula solitary, on peduncles
1–3 cm
long; hairs on achenes uncinate with bifid tips; creeping herb;
Madagascar
.......... ......................................................................
G. gossypina
8a. Capitula (at least the majority of them) in groups of 2–8, on short peduncles <
5 mm
long; hairs on achenes simple, not uncinate; erect or clambering herbs ........................ 9
9. Leaves all shallowly dentate with teeth to
3 mm
deep; phyllaries 1.2–1.5(–2) ×
0.3– 0.4 mm
; pappus bristles of outer florets in number 0–3(–5), of inner florets 4–9(–12); widespread in West and Central Africa ...
G. grangeoides
9a. Leaves, at least the lower ones, deeply pinnatilobed to pinnatifid; phyllaries 1.2 –2.5 ×
0.5–1.2 mm
; pappus bristles of outer florets in number 6– 16, of inner florets 12–18; endemic to west-central
Gabon
.................
G. ogoouensis