Pygommatius Scarbrough & Marascia, a valid ommatiine genus (Diptera: Asilidae) with new combinations and new species from the Philippines Author Scarbrough, A. G. Author Hill, H. text Zootaxa 2005 1055 1 22 journal article 10.5281/zenodo.170082 4f262a56-4eb7-45db-9462-9ecc42350767 1175­5326 170082 Pygommatius montanus Scarbrough & Hill , sp. nov. Figs. 30–38 Male. Brown, pleuron mostly and wide corners of most tergites yellow. Body length 10.6–11.9 mm ; wing length 8.5 mm . Head . Face gray, with yellow tint, vestiture mostly white, one or two thin brown bristles; mystax with several stout white bristles; setae sparse, scattered; FHWR 1.0: 9.6–1.0:11.6. Palpus brown, brown setose. Proboscis black, base yellow ventrally, yellow setose. Antenna brownish­yellow to brown, pedicel darkest; brown setose; FWLR 1.0:2.1–1.0:2.4. Frons contrastingly brownish­yellow tomentose, brown medially; brown setose. Ocellar tubercle brown setose, posterior setae as long as three antennal segments combined. Occiput with three or four brown postocular bristles dorsally, proclinate with longest just forward of posterior margin of eye. Thorax : Pronotum yellow to brown, yellow laterally. Mesonotum black, postalar corner reddish­yellow to yellow; setae sparse, mostly brown, thin; two dorsocentral and four lateral bristles, most black, one dorsocentral bristle yellow. Scutellum brown, yellowish­gray tomentose with scattered pale setae and two marginal bristles. Mediotergite black, anatergite reddish­yellow to yellow. Pleuron mostly yellow to reddishyellow; anepisternum, katepisternum, and meron partly black; tomentum mostly yellowish­gray, slightly brown anteriorly; vestiture yellow. Wing : Microtrichia present, dense from apex to crossvein r­m and apical 2/3 posteriorly. Cell r1 rounded apically. Cell r4 with base beyond apex of cell d, sides parallel, veins R4 and R5 divergent at wing margin. Crossvein r­m at apical 1/3 of cell d slightly shorter than CuA1. Cell m1 contrastingly wide beyond narrow base. Cell m3 short. Leg : Coxae mostly yellow; narrow apex of middle coxa and hind coxa anteriorly brown with brown setae anteriorly. Fore and middle trochanters yellow to brownishyellow; hind trochanter brown. Fore and middle femora yellow; hind femur with narrow brown band from middle to about apical 1/3. Fore femur with ventral row of stout yellow bristles. Middle femur with thin, mostly brown, anteroventral setae and stout, yellow, posteroventral bristles. Hind femur with sparse, thin, brown anteroventral setae; basal 1/3 with only three yellow setae, longest 1/2 as long as hind femur; abundant brown bristly setae posteriorly; HFWLR 1.0:4.6–1.0:5.0. Fore and middle tibiae entirely yellow with unusually long thin mostly yellow bristles. Hind tibia brown on apical 1/3. Basal tarsomere of fore and middle tarsi mostly yellow, narrow apex brown; bristles long, thin, usually brown; fore tarsus with one yellow bristle. Abdomen : Brown with wide apical corners and wide to narrow sides of tergites and apical margins of most sternites yellow; basal sternites with wide margins yellow. Apical margin of tergite 6 slightly produced medially. Tergites mostly brown setose, mostly short, yellow in light areas, often long; sternites 3–5 with stout, erect yellow bristles. Terminalia ( Figs. 30–34 ): Epandrium with dorsal branch broadly triangular, three short digits apically; middle branch abruptly capitate, apical margin evenly curved in dorsal and ventral views; ventral branch, short, wide and forked apically. Gonostylus narrowed apically, wider basally. Gonocoxite with deep notch along inner margin. Aedeagus wide apically, sheath hoodl­ike dorsally; ejaculatory apodeme wide with flange laterally. Hypandrium with transverse cluster of bristles apically. FIGURES 30–38. Pygommatius montanus Scarbrough , terminalia. Male, 30–32. Lateral, dorsal, and ventral views; 33. Gonostylus, lateral view; 34. Aedeagus lateral view. Female, 35–36. Dorsal and ventral views; 37. Spermatheca; 38. Genital fork. Female. Differs from male as follows. Body length 10.0– 11.4 mm ; wing length 8.1–9.4 mm ; FHWR 1.0:11.1–1.0:12.6; FWLR 1.0:1.8–1.0:2.0; cell m1WR 1.0:3.1:3.0– 1.0:4.0:3.8; HFWLR 1.0:6.2–1.0:6.3. Head : Face with six to eight long brown bristles. Antenna brown to reddish­brown; four or five postocular bristles brown. Thorax : Pronotum black. Dorsocentral bristles black. Pleuron mostly yellowish­red; katatergite with mostly brown bristles. Leg . Fore coxa with brown setae apically. Femora ventrally with setae and bristles thin and long. Middle femur with posteroventral bristles mostly brown, much thinner and longer than in male. Hind femur with all ventral bristles stout, and most black. Tibiae with mostly and tarsi with only black bristles. Abdomen : Apical margin of tergite 6 uniform, not produced medially; apical sternites with only scattered, thin, black setae. Sternite 8 with apical margin produced, broadly rounded medially and stout black bristles laterally. Terminalia ( Figs. 35–38 ): Segment 8 with unusually stout bristles laterally. Tergite 9 membranous dorsally. Sternite 8 long, strongly produced apically, apical margin rounded. Spermatheca bulbous basally, short, recurved. Genital fork with wide lateral and basal sclerites. Specimens examined. PHILIPPINES : Holotype ɗ, Mountain Prov. Abatan, Bugules, 60 km S of Bontoc, 1800–2000 m , 22–31.v.1964 ~ H. R. Torrevillas (BPBM #15,575). Paratypes , 2 Ψ, Camarines Sur: Mt. Isarog, 20 km E. of Naga, 500–600 m , 9.iv.1963 , light trap ~ H. R. Torrevillas (BPBM); 1 ɗ, Luzon, 2 km W of Santa Fe, Nueva Viscaya, 11.iv.1968 ~ D. E. Hardy (BPBM). Etymology : Latin montanus [feminine], referring to the type locality (mountains). Remarks. Pygommatius montanus is most similar to P. iriga , but is distinguished by the combined characters of the terminalia ( Figs. 30–38 ). The male also differs by the stout yellow bristles below the fore and middle femora, and the female by the strongly produced apical margin of sternite 8. Pygommatius montanus will run to O. hocus in Oldroyd’s key (1972) but can be distinguished by the shape of the epandrium.