Cladistic analysis of Rheochlus and related genera, with description of a new species (Diptera: Chironomidae: Podonominae) Author Siri, Augusto ILPLA (Instituto de Limnología ‘ Dr. Raúl A. Ringuelet’) CONICET-CCT-La Plata / UNLP. Bv. 120 y 62, La Plata (1900), Buenos Aires, Argentina; e-mail: augusto @ ilpla. edu. ar Author Brodin, Yngve The Swedish Museum of Natural History, Box 50007, SE- 104 05 Stockholm, Sweden; e-mail: yngve. brodin @ nrm. se Author Ilpla Author Conicet-Cct Author Unlp text Acta Entomologica Musei Nationalis Pragae 2014 2014-04-30 54 1 361 375 journal article 10.5281/zenodo.5299009 0374-1036 5299009 C954D5F0-070E-4A2B-93C7-FB1B49745B64 Rheochlus insignis Brundin, 1966 ( Figs 10–14 ) Type material examined. PARATYPES : 2 adult JJand 2 ♀♀ , “ CHILE / Cautín province / Río Llancahue at lago Pellaifa / 20-i-1958 / J. Illies col” ( SMNH ). Additional material examined. 1 adultJ and 1 adult ♀ , “ ARGENTINA / Río Negro province / Valle del Challhuaco / Mallín de los patos / 41°15’48.6’’S / 71°17’50.3’’W , 1020 m . a.s.l. / 10-i-2009 / Siri A. col / sweep net” ( MLPA ). Redescription. Male imago (n = 2–3) ( Figs 10–12 ). Total length 2.38–3.23 mm , total length / wing length 1.30–1.63. Head . Eyes hairy. Antenna with 14 flagellomeres ( Fig. 10 ). Ultimate flagellomere 150 long, penultimate 80 long. AR 0.43. Temporals 14, postorbitals 8 on each side. Clypeus with 2–4 setae. Palpomere lengths (I–V): 25; 20–25; 68–70; 45–48; 53. Tentorium 125 long. Thorax . Antepronotum with 3 setae. Dorsocentrals 19; acrostichals 20; prealars 12; supraalar 1; scutellars 8. Wing ( Fig. 11 ): length 1.63–1.98 mm ; width 0.54–0.69 mm ; L/ W 2.81 –2.90. C extension 90 long. VR 0.98–0.99. Width of cell r 1 at the end of vein R 1 115–145 ; maximum width of cell r 5 150–195; r 1 /r 5 0.72–0.79. Brachiolum with 3–7 setae; squama with 9–21 setae. R with 19–44 setae, R 1 with 22–51 setae, R 4 +5 with 27–42 setae, M 1+2 with 32 (1) setae, Cu 1 with 19 (1) setae, An with 8 setae (1). Cells r 4+5 and m 1+2 setose; remaining cells bare. Legs . Spur of front tibia 48–55 long; spurs of mid tibia 33–45 and 25–28 long; of hind tibia 54–73 and 25–33 long. Apex of ti 1 48–52 wide; of ti 2 50 wide; of ti 3 50–56 wide. Comb with 10–12 spines. Lengths and proportions of legs in Table 3. Hypopygium ( Fig. 12 ). Tergite IX with 22 setae (1), laterosternite IX with 3 setae (1). Transverse sternapodeme 125 long. Gonocoxite 160–200 long; dorsal keel 0.78–0.82 as long as gonocoxite. Gonostylus 95–120 long; basal portion of gonostylus swollen to nearly spherical; apical lobe 50–63 long, apical seta (“t” seta) 13 long; subapical lobe 19–25 long. Seta “p” 88–90 long, 4–5 wide, L/W of seta “p” 18–22. The “x” and “y” setae discernible, the “x” proximal and the “y” distal to mid section of the apical lobe. HR 1.67, HV 2.03. Female imago (n = 2–3) ( Figs 13–14 ). Total length 2.24–2.40 mm . Wing length / profemur length 3.29–3.46. Head . Hairy eyes. Antenna with 13 flagellomeres ( Fig. 13 ). Last flagellomere 83–113 long, broader than the preceding ones; about as long as flagellomeres 10–12. AR 0.18–0.20. Temporal setae 24; postorbitals 8 on each side (1). Palpomere lengths (I–V): 22; 45; 80; 48; 63 (1). Tentorium 88 long. Thorax . Setation (1): Antepronotum with 4 setae. Dorsocentrals 27; acrostichals 20; prealars 5; supraalar 1; scutellars 10. Wing ( Fig. 14 ): length 1.78–2.21 mm ; width 0.68–0.92 mm ; L/ W 2.40 –2.60. C extension 63–88 long. VR 0.97–1.02. Width of cell r 1 at the end of vein R 1 130–180 ; maximum width of cell r 5 160–230; r 1 /r 5 0.78–0.81. Brachiolum with 3–7 setae; squama with 4–7 setae. R with 25–41 setae, R 1 with 33–70 setae, R 4 +5 with 73–135 setae, M 1+2 with 39–83 setae, Cu 1 with 24–45 setae, An with 24 (1) setae. Almost all cells setose . Legs . Spur of front tibia 35–50 long; spurs of mid tibia 23–28 long and 28 long; of hind tibia 60–75 long and 23 long. Apex of ti 1 40–50 wide; of ti 2 40 (1) wide; of ti 3 45–60 wide. Comb with 10–11 spines. Lengths and proportions of legs in Table 4. Genitalia . Cercus 70–75 long, broadly triangular; noto plus ramus 200 long. Seminal capsule 40–58 long. Sternite VIII with +65 (1) setae. Tergite IX with 34 (1) setae. Segment X with 9 (1) setae on each side. Distribution. Chile (Cautín province), Argentina ( Rio Negro Province : Nahuel Huapi National Park).