On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea) Author Li, Yan-Da 0000-0002-9439-202X State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China ydli@pku.edu.cn Author Tihelka, Erik 0000-0002-5048-5355 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK & wn 20250 @ bristol. ac. uk; https: // orcid. org / 0000 - 0002 - 5048 - 5355 wn20250@bristol.ac.uk Author Dahan, Loïc 0000-0001-5680-0304 O. D. Taxonomy and Phylogeny - Entomology, Royal Belgian Institute of Natural Sciences, Vautierstreet 29, B- 1000 Brussels, Belgium ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X & loicdahan @ hotmail. com; https: // orcid. org / 0000 - 0001 - 5680 - 0304 loicdahan@hotmail.com Author Huang, Di-Ying 0000-0002-5637-4867 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867 dyhuang@nigpas.ac.cn Author Cai, Chen-Yang 0000-0002-9283-8323 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK & cycai @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 9283 - 8323 cycai@nigpas.ac.cn text Zootaxa 2021 2021-12-17 5082 3 223 244 journal article 2910 10.11646/zootaxa.5082.3.2 3997d831-ae29-42d0-ba99-cd20cc8b7fc3 1175-5326 5788224 EFB0E4A5-6B60-4B39-A651-1C6470FD7222 Archaenosodendron remotidens Li & Cai sp. nov. ( Figs 9–11 ) Material. Holotype , NIGP177614 . Etymology. The specific name refers to the relatively sparsely arranged denticles/stout setae on the outer edge of its protibia. Locality and horizon. Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District, Kachin State , Myanmar ; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian. Diagnosis. Medial antennomeres longer than wide ( Fig. 11C ). Lateral regions of mentum without longitudinally arranged larger punctures ( Fig. 11B ). Anterior angles of mentum not obviously produced ( Fig. 11B ). Antennal grooves on prosternum shorter than half-length of prosternum in front of coxae ( Fig. 11D ). Outer edge of protibia with about 14–16 relatively sparsely arranged denticles/stout setae, with the apical two distinctly stouter and more closely situated ( Fig. 11G ). Medial and lateral portions of prosternum not separated by a ridge ( Fig. 11D ). Description. Body elongated oval, about 3.6 mm long, 1.8 mm wide. Head prognathous. Eyes not strongly protuberant. Antennal insertions widely separated and concealed from above. Antennal grooves on head well developed. Frontoclypeal suture absent. Antennae 11-segmented; antennomere 2 about twice as long as wide; antennomere 3 about as long as but slender than 2; antennomeres 3–8 progressively shorter and wider; antennomeres 9–11 enlarged, forming a relatively loose club. Mandibles broad, unidentate. Maxilla with setose galea and lacinia; maxillary palps 4-segmented; palpomeres 1 and 4 longer than 2 and 3. Mentum with lateral edges subparallel in the basal half, then converging anteriorly; surface without clearly visible punctures; anterior angles not produced. Apical labial palpomere seemingly enlarged. Pronotal disc subtrapezoidal, widest at base; anterior margin with shallow emarginations laterally, slightly protruding medially. Prosternum in front coxae transverse, not extremely shortened; medial and lateral portions of prosternum not separated by a ridge; antennal grooves on prosternum shorter than half-length of prosternum in front of coxae; prosternal process relatively slender. Procoxae strongly transverse, well separated. Procoxal cavities broadly open posteriorly. FIGURE 8. Details of Archaenosodendron explanatum Li & Cai sp. nov. , holotype, NIGP177613, under confocal microscopy. A , Head, dorsal view. B , Head, ventral view. C , Head, lateral view. D , Prothorax, ventral view, showing the incomplete antennal groove on prosternum. E–G , Fore legs. H , Mid leg. I , Hind leg. Abbreviations: an1, antennomere 1; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; v1,2, ventrites 1,2. Scale bars: 200 μm. FIGURE 9. General habitus of Archaenosodendron remotidens Li & Cai sp. nov. , holotype, NIGP177614, under incident light. A , Dorsal view. B , Ventral view. Scale bars: 1 mm. Scutellar shield triangular, posteriorly acute. Elytra widest at anterior third; anterior corners weakly obtuse. Mesoventrite short, anteriorly with well-developed mesoventral cavity and procoxal rests. Mesocoxal cavities widely separated. Metaventrite transverse, flattened. Metacoxae narrowly separated. Femora with groove for receiving tibia. Tibiae expanded distally, with two apical spurs; outer edge of protibia with relatively sparsely arranged denticles/stout setae, with the apical two distinctly stouter and more closely situated. Tarsi 5-5-5; tarsomeres slender, not distinctly lobed; tarsomeres 1–4 ventrally with a cluster of long hairs at apex. Pretarsal claws simple. Abdomen with five ventrites. Ventrites 1 and 2 with a pair of impressions for receiving hind legs. Anterior margin of ventrites smooth, without notches.