An annotated catalog of Evanioidea (Insecta: Hymenoptera) from China Author Li, Yicheng College of Agricultural and Biotechnology, Hunan University of Humanities, Science and Technology, Loudi 417000, China. Author He, Yanbiao College of Agricultural and Biotechnology, Hunan University of Humanities, Science and Technology, Loudi 417000, China. & Loudi Institute of Agricultural Science, Loudi 417000, China. Author Chen, Huayan Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China. text Zootaxa 2024 2024-06-11 5468 1 75 111 http://dx.doi.org/10.11646/zootaxa.5468.1.3 journal article 298600 10.11646/zootaxa.5468.1.3 4c8198d6-b82c-4ff2-96f9-912012d39e7b 1175-5326 11614335 07BEBC41-572F-42E7-9B98-83A88467CF9C 5.1 Genus Gasteruption Latreille, 1796 Gasteruption : Latreille, 1796: 113–114 [descr., type species: Ichneumon assectator Linnaeus, 1758 designated by Latreille, 1810]; Crosskey, 1962: 381 [taxonomic history and complete list of synonyms], 383 [figs. 4–14, venation in some Gasteruption ]; Jennings & Austin, 2002: 743 [diag., comments, distribution and biology]; Jennings & Austin, 2004: 191–192 [host relationships of Gasteruption ]; Zhao et al ., 2012: 7–8 [descr.]; van Achterberg 2013: 59 [illustrated key for the Netherlands ]; van Achterberg & Talebi, 2014: 10 [illustrated key for Iran and Turkey ]; Tan et al ., 2016: 53–54 [genus taxonomy history, key to species Gastruption from Palaearctic China ]; Tan et al ., 2021: 3–4 [remarks, key to species Gastruption from Palaearctic China ]. Rhydinofoenus : Bradley, 1909: 38 [descr., type-species: Rhydinofoenus kaweahensis Bradley, 1909 by original designation]. Dolichofoenus : Kieffer, 1910: 77 [descr., type-species: Foenus raphidioides Westwood, 1851 by original designation]. Trichofoenus : Kieffer, 1910: 77 [descr., type-species: Gasteruption merceti Kieffer, 1904 by original designation]. Number of valid species from China : 41. (1) Gasteruption abeillei Kieffer, 1912 Gasteruption abeillei Kieffer, 1912: 228 [in key], 231[in key], 251[diag.]; Tan et al ., 2021: 10 [in key], 32 [in key], 37–38 [figs. 1–12, 3♂ 3♀ , NWUX , RMNH ]. Trichofoenus breviterebrae : Watanabe, 1934: 285 ; Synonymized with G. assectator (Linnaeus) by Pagliano & Scaramozzino (2000) and with G. boreale (Thomson) by Tan et al . (2016) and Johansson & van Achterberg (2016) . Distribution: China ( Shaanxi ), Russia , Europe ( Kieffer 1912 ; Tan et al . 2021 ). Biology. Host unknown. (2) Gasteruption amoyense Pasteels, 1958 Gasteruption amoyense : Pasteels, 1958: 178–179 [descr., fig. 4, USNM , ♀ ]; Zhao et al ., 2012: 13 [in key, female], 16 [in key, male], 17–19 [descr., figs. 2–9; 1♀ , ZJU ; 3♀♀ , CSCS ]; Tan et al ., 2021: 21 [in key], 28 [in key], 38–42[figs. 13–26, 3♀ , NWUX ]. Gasteruption curiosum : Pasteels, 1958: 177–178 [descr., ♀ , NHMUK , Type Hym. 3a.365, fig. 3.] . Distribution: China ( Shaanxi , Zhejiang , Fujian , Hunan , Hongkong ) ( Pasteels 1958 ; Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (3) Gasteruption angulatum Zhao, van Achterberg & Xu, 2012 Gasteruption angulatum : Zhao et al ., 2012: 12 [in key, female], 16 [in key, male], 19–22 [descr., figs. 10–23, holotype , ♀ , ZJU ; paratype : 1♀ , ZJU ; 1♂ , SCAU ; 2♀ , SCAU ; 2♀ 1♂ , ZJU ; 1♀ , RMNH , 1♂ , ZJU ; 1♂ , SCAU ]. Tan et al ., 2016: 66 [in key], 76 [in key], 85 [figs. 1–6, 1♂ , NWUX ]; Tan et al ., 2021: 17 [in key], 28 [in key], 42[figs. 27–37, 1♂ , NWUX ]. Distribution: China ( Shaanxi , Henan , Hubei , Zhejiang ) ( Zhao et al . 2012 ; Tan et al . 2016 ; Tan et al . 2021 ). Biology. Host unknown. (4) Gasteruption assectator ( Linnaeus, 1758 ) Ichneumon assectator : Linnaeus, 1758: 566 [descr.]; Foenus assectator : Fabricius, 1798: 240 [descr.]; Westwood, 1843: 255 [note on taxonomy]. Gasteruption assectator : Schletterer, 1885: 276–277 [descr.], 316–317 [in catalog]; Dalla Torre, 1902: 1063 [in catalog]; Kieffer, 1912: 256 [descr., fig. 63]; Hedicke, 1939: 5 [in catalog.]; Townes, 1950: 121 [in key], 123–128 [descr.]; Madl, 1989b: 159 [descr.], 1989c: 41 [in key]; Kofler & Madl, 1990: 320 [in the catalog]; Wall, 1994: 150–151 [descr.]; Smith, 1996: 492 [in key]; Pagliano & Scaramozzino, 2000: 11 [in key, female], 19 [in key, male]; Zhao et al ., 2012: 10 [in key, female], 16 [in key, male], 23–27 [descr., figs. 24–37, numerous specimens deposited in ZJU and CSCS ]; Tan et al ., 2021: 9 [in key], 31 [in key], 42–44[notes]. Foenus borealis : Thomson, 1883: 849 [descr.]; Hedicke, 1939: 7 [in catalog]. Synonymized with G. assectator Linnaeus by Schletterer, 1889 . Gasteruption boreale : Schletterer, 1885: 303–304 [descr.]. Foenus fumipennis : Thomson, 1883: 848 [descr.]; Hedicke, 1939: 7 [in catalog]; Wall, 1994: 148 [synonymy with G. assectator ]. Synonymized with G. assectator Linnaeus by Schletterer, 1885 . Foenus nigritarsis : Thomson, 1883: 849 [descr.]; Schletterer, 1889: 398 [taxonomic history]; Hedicke, 1939: 7 [in catalog]; Wall, 1994: 149 [synonymy with G. assectator ]. Synonymized with G. assectator Linnaeus by Schletterer, 1889 . Gasteruption nigritarse : Schletterer, 1885: 310 [descr.]. Gasteruption brevicauda : Kieffer, 1904: 648 [descr.], 1912: 259 [descr.]; Hedicke, 1939: 8 [in catalog]; Madl, 1987a: 401 [in key]; Wall, 1994: 148 [synonymy with G. assectator ]. Synonymized with G. assectator Linnaeus by Madl, 1987b . Gasteruption abeillei : Kieffer, 1912: 228 [in key, female], 231 [in key, male], 251 [descr.]; Hedicke, 1939: 5 [in catalog]; Wall, 1994: 148 [synonymy with G. assectator ]. Synonymized with G. assectator Linnaeus by Madl, 1989b . Trichofoenus breviterebrae : Watanabe, 1934: 285 [descr.]; Hedicke, 1939: 45 [in catalog]. Gasteruption rugulosum : Malyshev, 1965: 245 [descr.]. Gasteruption margotae : Madl, 1987b:225 [descr.]; Wall, 1994:149 [synonymy with G.assectator ].Synonymized by with G. assectator Linnaeus by Madl, 1990 . Distribution: China ( Heilongjiang , Jilin , Inner Mongolia , Xinjiang , Beijing , Hebei , Henan , Shanxi , Ningxia , Qinghai , Hubei , Tibet , Hunan , Sichuan ), Russia , Japan , Europe ( Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (5) Gasteruption assectoides Zhao, van Achterberg & Xu, 2012 Gasteruption assectoides : Zhao et al ., 2012: 11 [in key, female], 28–30 [descr., figs. 38–46, holotype , ♀ , CSCS ] . Distribution: China ( Hubei ) ( Zhao et al . 2012 ). Biology. Host unknown. (6) Gasteruption bicoloratum Tan & van Achterberg, 2016 Gasteruption bicoloratum : Tan et al ., 2016: 61 [in key], 78 [in key], 86–90, [figs. 7–22, holotype , ♀ , NWUX , paratype , 4♀♀ 3♂♂ , NWUX and RMNH .]; Tan et al ., 2021: 10 [in key], 31 [in key], 44[ 1♂ 1♀ , NWUX ] Distribution: China ( Gansu , Shaanxi ), Russia ( Tan et al . 2016 ; Tan et al . 2021 ). Biology. Host unknown. (7) Gasteruption bimaculatum Pasteels, 1958 Gasteruption bimaculatum : Pasteels, 1958: 191–192 , [fig. 16, holotype , ♂ , NHRS -HEVA000001977, paratype , 2♀♀ , USNM ]; Zhao et al ., 2012: 10 [in key, female], 14 [in key, male], 30–35 [descr., figs. 47–62, paratype , 1♂ 3♀♀ , CSCS , 1♂ 7♀♀ , ZJU , 1♀ , RMNH , 2♀♀ , SCAU ]; Tan et al ., 2021: 11 [in key], 32 [in key], 44–45 [figs. 38–56; 2♂ 3♀ , NWUX , 1♀ , RMNH ]. Gasteruption obscuripenne : Pasteels, 1958: 189–190 [descr., fig. 14]. Distribution: China ( Shaanxi , Henan , Fujian , Hainan , Guangxi , Yunnan , Tibet ), Burma ( Pasteels 1958 ; Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (8) Gasteruption birmanense Pasteels, 1958 Gasteruption birmanense : Pasteels, 1958: 199 [ holotype , ♀ , NHRS -HEVA 000001978; paratype , 1♂ , NHRS - HEVA000001979], fig. 22; Zhao et al ., 2012: 13 [in key, female], 16 [in key, male], 35–38 [descr., figs. 63–70], [ 1♀ , ZJU , 1♀ , CSCS ]. Distribution: China ( Hunan , Sichuan ), Burma ( Pasteels 1958 ; Zhao et al . 2012 ). Biology. Host unknown. (9) Gasteruption boreale ( Thomson, 1883 ) Foenus borealis ; Thomson, 1883: 849 [descr.]. Synonymized with G. assectator (Linnaeus) by Schletterer (1889) and with G. minutum (Tournier) by van Achterberg & Talebi (2014) . Gasteruption boreale : Schletterer, 1885: 303 [descr.]. Tan et al ., 2016: 59 [in key, female], 79 [in key], 90–91 [diag., figs. 23–32, holotype of G. breviterebrae , ♀ , ECHU , holotype of Trichofoenus breviterebrae ; paratype , 1♂ , ECHU ]. Trichofoenus breviterebrae ; Watanabe, 1934: 285 [descr.]. Synonymized with G. assectator (Linnaeus) by Pagliano & Scaramozzino (2000) and with G. boreale (Thomson) by Johansson & van Achterberg (2016) . Distribution: China ( Heilongjiang ), Russia ( Thomson 1883 ; Tan et al . 2016 ). Biology. Host unknown. (10) Gasteruption brevicuspis Kieffer, 1911 Gasteruption brevicuspis : Kieffer, 1911: 196 [descr.], 1912: 293–294 [descr.]; Tan et al ., 2021: 11 [in key], 31 [in key], 45–51 [figs. 57–79; 16♂ 30♀ , NWUX , 1♂ 5♀ , RMNH ]. Gasteruption terebrelligerum : Enderlein, 1913: 323–324 [descr., holotype , ♀ , SDEI ]; Hedicke, 1939: 28 [in catalog]; Pasteels, 1958: 189 [descr.]; Zhao et al ., 2012: 10 [in key, female], 16 [in key, male], 97–100 [descr., figs. 273–285; 1♀ 3♂♂ , SCAU, numerous specimens from Fujian , Shanxi , Hunan and Yunnan , CSCS ]. Distribution: China ( Gansu , Shaanxi , Shanxi , Hubei , Fujian , Taiwan , Hunan, Yunnan) ( Enderlein 1913 ; Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (11) Gasteruption coloratum Zhao, van Achterberg & Xu, 2012 Gasteruption coloratum : Zhao et al ., 2012: 11 [in key, female], 16 [in key, male], 38–40 [descr., figs. 71–78, holotype , ♀ , ZJU ]; Tan et al ., 2021: 13 [in key], 33 [in key], 51–52 [figs. 80–92; 2♀ , NWUX ]. Distribution: China ( Xinjiang ) ( Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (12) Gasteruption corniculigerum Enderlein, 1913 Gasteruption corniculigerum : Enderlein, 1913: 322–323 [descr., paratype , 4♀ 2♂ , SDEI ]; Hedicke, 1939: 27 [in catalog]; Pasteels, 1958: 176–177 [descr.], fig. 1; Zhao et al ., 2012: 11 [in key, female], 14 [in key, male], 40–45 [descr., figs. 79–94, 9♀ 6♂ , ZJU , 1♀ 1♂ , NMNS , 5♀ 3♂ , TARI , 17♂ , SCAU ]; Tan et al ., 2021: 16 [in key], 23 [in key], 53–54 [figs. 93–106; 2♀ , NWUX, 1♀ , RMNH ]. Distribution: China ( Shaanxi , Zhejiang , Fujian , Taiwan , Hunan, Guangdong, Guangxi, Hainan, Guizhou) ( Enderlein 1913 ; Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host, Hylaeus perforata ( Guo et al . 2021 ) . (13) Gasteruption dilutum Semenov, 1892 Gasteryption dilutum : Semenov, 1892: 23–24 [ holotype , ♂ , ZISP ] . Gasteruption dilutum ; Szépligeti, 1903: 369 [in key]; Kieffer, 1912: 231 [in key], 271 [descr.]; Hedicke, 1939: 8 [in catalog]; Zhao et al ., 2012: 11 [in key, female], 16 [in key, male], 45–47 [descr., figs. 95–102]. Distribution: China ( Xinjiang ) ( Semenov 1892 ; Zhao et al . 2012 ). Biology. Host unknown. (14) Gasteruption dimidiatum Semenov, 1892 Gasteryption dimidiatum : Semenov, 1892: 22–23 [descr., holotype , ♀ , ZISP, 79396]; Kieffer, 1912: 229 [in key], 263–264 [descr.]; Hedicke, 1939: 9 [in catalog]; Zhao et al ., 2012: 10 [in key, female], 14 [in key, male], 48–50 [descr., figs. 103–110, paratype , 4♀♀ , ZJU , 1♀ , RMNH ]. Distribution: China ( Xinjiang ), Uzbekistan ( Semenov 1892 ; Zhao et al . 2012 ). Biology. Host unknown. (15) Gasteruption formilis Alekseev, 1995 Gasteruption formilis : Alekseev, 1995: 42 [in key only, holotype information unavailable]; Zhao et al ., 2012: 10 [in key, female], 15 [in key, male], 51–54 [descr., figs. 111–123; 3♂♂ 1♀ , ZJU ]. Distribution: China ( Gansu , Xinjiang ) ( Alekseev 1995 ; Zhao et al . 2012 ). Biology. Host unknown. (16) Gasteruption formosanum Enderlein, 1913 Gasteruption formosanum : Enderlein, 1913: 326 [descr., holotype , ♂ , SDEI ]; Hedicke, 1939: 27 [in catalog]; Pasteels, 1958: 188–189 [descr.], fig. 13; Zhao et al ., 2012: 9 [in key, female], 14 [in key, male], 54–57 [descr., 1♀ 3♂ , RMNH , 2♀ , ZJU , 2♀ , RMNH , figs. 124–141]. Distribution: China ( Zhejiang , Fujian , Hunan , Taiwan) ( Enderlein 1913 ; Zhao et al . 2012 ). Biology. Host unknown. (17) Gasteruption granulatum Tan & van Achterberg, 2021 Gasteruption granulatum : Tan et al ., 2021: 7 [in key], 29 [in key], 55–59, [descr., figs. 107–123, holotype, ♀ , NWUX , paratype, 1♀ , RMNH , 1♂ 1♀ , NWUX ]. Gasteruption parvicollarium : Zhao et al ., 2012: 68–72 [female]. Distribution: China ( Shaanxi ) ( Tan et al . 2021 ). Biology. Host unknown. (18) Gasteruption huangshii Tan & van Achterberg, 2016 Gasteruption huangshii : Tan et al ., 2016: 67 [in key], 75 [in key], 91–96, [figs. 33–48, holotype, ♀ , NWUX , paratype, 5♂♂ , NWUX and RMNH .] Distribution: China ( Shaanxi ) ( Tan et al . 2016 ). Biology. Host unknown, but this species was collected together with a Hylaeus sp. ( Tan et al . 2016 ). (19) Gasteruption japonicum Cameron, 1888 Gasteruption japonicum : Cameron, 1888: 134 [descr., holotype , ♀ , NHMUK ]; Zhao et al ., 2012: 13 [in key, female], 15 [in key, male], 58–61 [descr., 5♀ 2♂ , TARI , 9♀ , ZJU , 5♀ 1♂ , CAU , 2♀ , SCAU , figs. 142–152]; Tan et al ., 2016: 71 [in key, female], 84 [in key], 97–98 [notes., figs. 49–57, 2♀ , NWUX , RMNH ; 2♀ 1♂ , NWUX ]; Tan et al ., 2021: 22 [in key], 36 [in key], 59–60 [notes, 4♂ 22♀ , NWUX, 4♀ , RMNH ] Gasteruption sinense var. minus : Kieffer, 1924: 78 [descr., holotype , 1♀ 1♂ , TARI , probably lost or present in remants of Kieffer collection] . Distribution: China ( Heilongjiang , Jilin , Inner Mongolia , Shaanxi , Ningxia , Gansu , Xinjiang , Shanghai , Zhejiang , Fujian , Taiwan , Hubei, Hunan, Sichuan, Yunnan), Japan ( Cameron 1888 ; Kieffer 1924 ; Zhao et al . 2012 ; Tan et al . 2016 ; Tan et al . 2021 ). Biology. Host unknown. (20) Gasteruption rufescenticorne Enderlein, 1913 Gasteruption rufescenticorne : Enderlein, 1913: 324–325 [descr., lectotype , 1♀ , ZMIZ , 2♂ , SDEI , 1♂ , ZMIZ ]; Hedicke, 1939: 28 [in catalog]; Pasteels, 1958: 177 [descr., fig. 2]; Zhao et al ., 2012: 11 [in key, female], 14 [in key, male], 75–80 [descr., 5♀♀ 19♂♂ , ZJU , 1♀ , CAU , 1♂ , NMNS , 5♀♀ 5♂♂ , TARI , 3♀♀ 4♂♂ , SCAU , figs. 200–216]. Distribution: China ( Ningxia , Jiangsu , Shanghai , Zhejiang , Fujian , Taiwan , Hubei, Hunan, Guangxi, Hainan) ( Enderlein 1913 ; Zhao et al . 2012 ). Biology. Host unknown. (21) Gasteruption kexinae Tan & van Achterberg, 2021 Gasteruption rufescenticorne : Zhao et al ., 2012: 75–80 [ paralectotype ; not lectotype ]. Gasteruption kexinae : Tan et al ., 2021: 16 [in key], 23 [in key], 60–65 [descr., figs. 144–154, holotype, ♀ , NWUX , paratype, 3♀ 3♂ , NWUX , RMNH , 3♀ 10♂ , ZJU ]. Distribution: China ( Ningxia , Jiangsu , Shanghai , Zhejiang , Fujian , Taiwan , Hubei, Hunan, Guangxi) ( Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (22) Gasteruption latitibia Zhao, van Achterberg & Xu, 2012 Gasteruption latitibia : Zhao et al ., 2012: 9 [in key, female], 16 [in key, male], 62–65 [descr.; holotype , ♀ , ZJU ; paratypes , 2♀♀ 1♂ , ZJU , 1♀ 1♂ , SCAU , figs. 153–165]; Tan et al ., 2021: 8 [in key], 29 [in key], 65–66 [descr., figs. 144–154, 6♂ 20♀ , NWUX , 5♀ , RMNH ]. Distribution: China ( Shaanxi , Hubei , Hunan , Zhejiang , Fujian , Guizhou ) ( Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (23) Gasteruption minutum ( Tournier, 1877 ) Foenus minutus : Tournier, 1877 : ix [descr.]; Schletterer, 1889: 398 [descr.] (as synonym of G. assectator (Linnaeus)) . Gasteruption minutum : Kieffer, 1912: 257 [descr.]; Wall, 1994: 159 [descr.]; Pagliano & Scaramozzino, 2000: 11 [in key], 19 [in key], 30 [descr.]; Tan et al ., 2021: 5 [in key], 24 [in key], 66–71 [notes, figs. 154–173, 5♀ 1♂ , NWUX, RMNH ]. Foenus longigena : Thomson, 1883: 849 [descr.]. Synonymized with G. minutum (Tournier) by Ferrière (1946). Gasteruption longigena : Schletterer, 1889: 399 [descr.]; Kieffer, 1912: 270 [descr.]; Wall, 1994: 149 [descr.]; Hellén, 1950: 4 [as G. “ longiserra ” and as synonym of G. minutum (Tournier) ]. Gasteruption oriplanum : Kieffer, 1911: 210 [descr.]; Zhao et al ., 2012: 8 [in key], 15 [in key], 65–68 [descr., figs. 166–173, 3♂ , ZJU, RMNH, 5♀ , CSCS ]. Synonymized with G. minutum by Tan et al . (2021) . Distribution: China ( Tibet , Shaanxi ), South Palaearctic, Central Europe ( Tournier 1877 ; Kieffer 1912 ; Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (24) Gasteruption oriplanum Kieffer, 1911 Gasteruption oriplanum : Kieffer, 1911: 210 [descr., holotype , ♂ , NHMUK ]; Hedicke, 1939: 27 [in catalog]; Zhao et al ., 2012: 8 [in key, female], 15 [in key, male], 65–68 [descr., 3♂♂ , ZJU and RMNH , 1♀ , CSCS , figs. 166–173]. Distribution: China ( Tibet ) ( Kieffer 1911 ; Zhao et al . 2012 ). Biology. Host unknown. (25) Gasteruption nigritarsis ( Thomson, 1883 ) Foenus nigritarsis : Thomson, 1883: 849 [descr.]; Schletterer, 1889: 398 [descr.]; Wall, 1994: 149 [descr.]. Synonymized with G. assectator (Linnaeus) by Schletterer (1889) . Gasteruption nigritarse : Schletterer, 1885: 310 [descr.]; Johansson & van Achterberg, 2016: 84–86 [descr.]; Tan et al ., 2021: 9 [in key], 31 [in key], 71–75 [notes, figs. 174–194, 6♀ , NWUX, 5♀ 1♂ , RMNH ]. Distribution: China ( Gansu , Shaanxi ), Europe ( Thomson 1883 ; Schletterer 1885 ; Tan et al . 2021 ). (26) Gasteruption oshimense Watanabe, 1934 Gasteruption tournieri : Schletterer, 1885: 287 [descr., holotype , ♀ ]; Dalla Torre, 1902: 1075 [in the catalog]; Szépligeti, 1903: 368 [in key]; Kieffer, 1912: 227 [in key, female], 230 [in key, male], 247 [descr.]; Hedicke, 1939: 25 [in catalog]; Crosskey, 1951: 292–294 [descr., fig. 9]; Madl, 1987a: 404 [diag.]; Kofler & Madl, 1990: 322 [in catalog]; Wall, 1994: 164 [descr.]; Pagliano & Scaramozzino, 2000: 15 [in key, female], 17 [in key, male], 33–34 [descr.]; Turrisi, 2004: 84 [in catalog]; Zhao et al ., 2012: 11 [in key, female], 13 [in key, male], 103–108 [descr., figs. 295–308; 1♀ , CSCS , 7♀♀ 1♂ , ZJU , 5♀♀ , SCAU ]. Gasteruption austriacum : Schletterer, 1885: 277 [descr., holotype , ♂ ]; Hedicke, 1939: 26 [in catalog]; Wall, 1994: 148 [synonymy with G. tournieri ]; synonymized with G. tournieri Schletterer by Schletterer, 1889 . Gasteruption nitidum : Schletterer, 1885: 281–282 [descr., holotype , ♂ ]; Hedicke, 1939: 26 [in catalog]; Wall, 1994: 149 [synonymy with G. tournieri ]; synonymized with G. tournieri Schletterer by Schletterer, 1889 . Gasteruption oshimense Watanabe, 1934: 283–284 . Tan et al ., 2016: 65 [in key], 71 [in key], 99–100 [notes, figs. 58–67, 2♀ 3♂ , NWUX , 3♀ 6♂ , NWUX , RMNH , G. tournieri by Zhao et al ., 2012 are included under Gasteruption oshimense ]; Tan et al ., 2021: 15 [in key], 23 [in key], 75–78 [notes, figs. 195–212, 20♀ 6♂ , NWUX, 8♀ , RMNH ]. Distribution: China ( Gansu , Jilin , Shaanxi , Henan , Hunan , Guizhou ), Japan , South and Central Europe, up to England , Netherlands and Germany ( Watanabe 1934 ; Zhao et al . 2012 ; Tan et al . 2016 ; Tan et al . 2021 ). Biology. Host unknown. (27) Gasteruption pannuceum Tan & van Achterberg, 2016 Gasteruption pannuceum : Tan et al ., 2016: 63 [in key], 76 [in key], 101–102, [descr., figs. 68–77, holotype , ♀ , NWUX ] Distribution: China ( Shaanxi ) ( Tan et al . 2016 ). Biology. Host unknown. (28) Gasteruption parvicollarium Enderlein, 1913 Gasteruption parvicollarium : Enderlein, 1913: 323–324 [descr., holotype , ♂ , SDEI ]; Hedicke, 1939: 27 [in catalog]; Pasteels, 1958: 183 [descr., fig. 8]; Zhao et al ., 2012: 9 [in key, female], 15 [in key, male], 68–72 [descr., figs. 174–190, 8♀♀ 4♂♂ , ZJU , 1♂ , CAU , 1♀ , RMNH , 10♀♀ 20♂♂ , TARI , 3♀♀ , SCAU ]; Tan et al ., 2021: 7 [in key], 27 [in key], 78–82 [notes, figs. 213–231, 9♀ 4♂ , NWUX, 4♀ , RMNH ]. Distribution: China ( Shaanxi , Fujian , Taiwan) ( Enderlein 1913 ; Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (29) Gasteruption pedion Tan & van Achterberg, 2021 Gasteruption pedion : Tan et al ., 2021: 20 [in key], 27 [in key], 82–86, [descr., figs. 232–252, holotype , ♀ , NWUX , paratype , 2♂ 3♀ , RMNH , NWUX , 1♂ 1♀ , SCAU ]. Distribution: China ( Jiangsu , Shaanxi , Yunnan ) ( Tan et al . 2021 ). Biology. Host unknown. (30) Gasteruption poecilothecum Kieffer, 1911 Gasteruption poecilothecus : Kieffer, 1911: 205 [descr., holotype , ♀ , NHMUK ]; Zhao et al ., 2012: 13 [in key, female], 16 [in key, male], 73–75 [descr., 2♀♀ , ZJU , 1♀ , RMNH , 1♀ , CSCS , figs. 191–199]. Distribution: China ( Heilongjiang , Jilin , Xinjiang , Hebei ) ( Kieffer 1911 ; Zhao et al . 2012 ). Biology. Host unknown. (31) Gasteruption reductum Tan & van Achterberg, 2021 Gasteruption pedion : Tan et al ., 2021: 19 [in key], 27 [in key], 86–91 [descr., figs. 253–271, holotype, ♀ , NWUX , paratype, 1♂ 5♀ , RMNH , NWUX , 1♀ , SCAU ]. Distribution: China ( Shaanxi , Hebei ) ( Tan et al . 2021 ). Biology. Host unknown. (32) Gasteruption shengi Tan & van Achterberg, 2016 Gasteruption shengi : Tan et al ., 2016: 63 [in key], 82 [in key], 104–108, [descr., figs. 78–94, holotype, ♀ , NWUX , paratype, 1♀ 1♂ , RMNH , 1♂ , NWUX .] Distribution: China ( Ningxia , Inner Mongolia ) ( Tan et al . 2016 ). Biology. Host unknown. (33) Gasteruption sinarum Kieffer, 1911 Gasteruption sinarum : Kieffer, 1911: 205–206 [descr., holotype , ♀ , NHMUK ]; Kieffer, 1912: 229 [in key], 264 [descr.]; Hedicke, 1939: 21 [in catalog]; Zhao et al ., 2012: 12 [in key, female], 16 [in key, male], 80–85 [descr., 21♀♀ 4♂♂ , ZJU , 6♀♀ 1♂ , CAU , 1♀ , RMNH , 28♀ 3♂♂ , TARI , 29♀♀ 5♂♂ , SCAU , 1♀ 1♂ , CSCS , figs. 217– 227]; Tan et al ., 2021: 13 [in key], 37 [in key], 91–92 [notes, figs. 272–291, 1♀ 1♂ , NWUX ]. Gasteruption sinense : Kieffer, 1924: 77–78 [descr., lectotype , ♀ , TARI, paralectotype , 1 ♀ , TARI ]; Hedicke, 1939: 21 [in catalog]. Distribution: China ( Liaoning , Inner Mongolia , Beijing , Tianjin , Ningxia , Shaanxi , Henan , Jiangsu , Shanghai , Shandong , Zhejiang , Anhui , Hubei , Hunan , Guangdong , Guangxi , Guizhou ) (Kiffer 1911; Kieffer 1924 ; Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (34) Gasteruption sinepunctatum Zhao, van Achterberg & Xu, 2012 Gasteruption sinepunctatum : Zhao et al ., 2012: 13 [in key, female], 15 [in key, male], 85–87 [descr.; holotype , ♀ , ZJU ; paratype , 3♀♀ 2♂♂ , TARI , 1♀ , CSCS ; figs. 228–240]; Tan et al ., 2016: 70 [in key], 75 [in key], 108–109 [note, figs. 95–104, 1♀ , NWUX ]; Tan et al ., 2021: 22 [in key], 27 [in key], 92–96 [notes, figs. 292–298, 5♀ 3♂ , NWUX ]. Distribution: China ( Jilin , Shaanxi , Zhejiang , Taiwan , Tibet) ( Zhao et al . 2012 ; Tan et al . 2016 ; Tan et al . 2021 ). Biology. Host unknown. (35) Gasteruption sinicola ( Kieffer, 1924 ) Trichofoenus sinicola : Kieffer, 1924: 79 [descr., holotype , probably lost]; Hedicke, 1939: 45 [in catalog]. Gasteruption sinicola : Zhao et al ., 2012: 9 [in key, female], 15 [in key, male], 88–91 [descr., figs. 241–251, 11♀♀ 10♂♂ , ZJU , 2♀♀ 3♂♂ , SCAU , 1♀ , RMNH , combination with Trichofoenus sinicola ]. Distribution: China ( Ningxia , Jiangsu , Fujian , Hunan , Guangdong , Hainan ) ( Kieffer 1924 ; Zhao et al . 2012 ). Biology. Host unknown. (36) Gasteruption strigosum Zhao, van Achterberg & Xu, 2012 Gasteruption strigosum : Zhao et al ., 2012: 10 [in key, female], 15 [in key, male], 91–94 [descr., figs. 252–264, holotype , ♀ , SCAU ; paratype , 2♀♀ 1♂ , ZJU , 1♀ 1♂ , CSCS ]. Distribution: China ( Fujian , Hainan , Yunnan ) ( Zhao et al . 2012 ). Biology. Host unknown. (37) Gasteruption subhamatum Pasteels, 1958 Gasteruption subhamatum : Pasteels, 1958: 206–207 [fig. 28; holotype , ♀ , USNM ]; Zhao et al ., 2012: 8 [in key, female], 14 [in key, male], 94–97 [descr., figs. 265–272; 2♀♀ , SCAU ]. Distribution: China ( Hainan ), Malaysia ( Pasteels 1958 ; Zhao et al . 2012 ). Biology. Host unknown. (38) Gasteruption subtile ( Thomson, 1883 ) Foenus subtilis : Thomson, 1883: 847 [descr.]. Gasteruption subtile : Schletterer, 1889: 425 [descr.]; Szépligeti, 1903: 370 [descr.]; Kieffer, 1912: 263 [descr.]; Pagliano & Scaramozzino, 2000: 13 [in key], 19 [in key], 33 [descr.]; Tan et al ., 2021: 20 [in key], 36 [in key], 96–98 [notes, figs. 299–307, 1♀ 1♂ , NWUX, 1♀ , NHMUK ]. Gasteruption kriechbaumeri : Schletterer, 1889: 384 [in key], 389[in key], 395–396 [descr.]; Szépligeti, 1903: 370 [descr.]; Kieffer, 1912: 267 [descr.]; Wall, 1994: 149 [descr.]. Synonymized with G. subtile (Thomson) by Madl (1989a) . Gasteruption sabulosum : Schletterer, 1889: 390 [in key], 396 [diag.]; Kieffer, 1912: 264 [descr.]; Wall, 1994: 149 [descr.]. Synonymized with G. kriechbaumeri Schletterer by Ferrière (1946) and with G. subtile (Thomson) by Madl (1989a) . Gasteruption poecilothecus : Kieffer, 1911: 205 [descr.]. Gasteruption poecilothecum : Zhao et al . 2012: 73–75 [descr.]; Tan et al . 2016: 69 [in key], 84 [in key]. Gasteruption rossicum : Kozlov, 1974: 76 [descr.]; Wall, 1994: 149 [descr.]. Synonymized with G. subtile (Thomson) by Kozlov (1974) . Distribution: China ( Shaanxi , Hebei , Heilongjiang , Inner Mongolia , Jilin , Xinjiang ); Europe (alpine-boreal); Mongolia ; Far East Russia ( Thomson 1883 ; Kieffer 1911 ; Zhao et al . 2012 ; Tan et al . 2021 ). Biology. Host unknown. (39) Gasteruption tonkinense Pasteels, 1958 Gasteruption tonkinense : Pasteels, 1958: 180 [descr., holotype , ♀ , MNHM ]; Zhao et al ., 2012: 12 [in key, female], 13 [in key, male], 100–103 [descr., figs. 286–294; 1♀ , SCAU, 1♀ , TARI , 5♀♀ , ZJU , 2♀♀ , CSCS ]. Distribution: China ( Shanghai , Zhejiang , Fujian , Guangxi , Guizhou ), Vietnam ( Pasteels 1958 ; Zhao et al . 2012 ). Biology. Host unknown. (40) Gasteruption transversiceps Pasteels, 1958 Gasteruption transversiceps : Pasteels, 1958: 208–209 [descr., holotype , ♀ , USNM ], fig. 30; Zhao et al ., 2012: 12 [in key, female], 16 [in key, male], 108–110 [descr., figs. 309–316, 1♀ , ZJU ]. Distribution: China ( Hainan ), Malaysia ( Pasteels 1958 ; Zhao et al . 2012 ). Biology. Host unknown. (41) Gasteruption varipes ( Westwood, 1851 ) Foenus varipes : Westwood, 1851: 220 [descr., holotype , ♂ , NHMUK ]. Gasteruption varipes : Schletterer, 1885: 289–290 [descr.], 325 [in catalog], 1890: 383 [in key], 434 [descr.]; Kieffer, 1912: 231 [in key], 269 [redescr.]; Hedicke, 1939: 26–27 [in catalog]; Pasteels, 1958: 186–188 [descr.]; Zhao et al ., 2012: 11 [in key, female], 14 [in key, male], 111–117 [descr., figs. 317–333; 7♂♂ , ZJU and RMNH , 11♀♀ 12♂♂ , ZJU , 1♀ , RMNH , 1♀ , TARI , 1♀ , CSCS ]. Distribution: China ( Fujian , Taiwan , Hainan, Yunnan) ( Westwood 1851 ; Zhao et al . 2012 ). Biology. Host unknown.