Sophrosynidae, a new family in the Lysianassoidea (Crustacea: Amphipoda) with a revision of the genus Sophrosyne Author Lowry, J. K. Author Stoddart, H. E. text Zootaxa 2010 2010-02-25 2370 1 1 35 https://biotaxa.org/Zootaxa/article/view/zootaxa.2370.1.1 journal article 10.11646/zootaxa.2370.1.1 1175-5326 5306939 Sophrosyne inverarae sp. nov. ( Figs10–12 ) Sophrosyne robertsoni . Norman, 1900: 196 (in part). — Moore, 1984: 37 (in part, part = S. moorei ). — Lincoln, 1979: 52 , fig. 17. — Kilgallen et al. , 2007: 1243 , fig. 1. Type material . HOLOTYPE , female, ovigerous (at least 5 eggs ), 6.5 mm , BMNH 1911:11:8:12449-12551, Inveraray , Loch Fyne , Scotland (approx. 56°13'N 5°4'W ), 1874 . PARATYPE , female, same locality [material recorded by Norman (1900) as S. robertsoni ]. Type locality. Inveraray , Loch Fyne , Scotland (approx. 56°13'N 5°4'W ) . Etymology . The specific name is a reference to the type locality. Diagnostic description . Head lateral cephalic lobe small, narrowly rounded; eyes apparently absent. Antenna 1 peduncular article 1 short, length 1.3 x breadth; accessory flagellum medium length, 0.5 x primary flagellum, article 1 short, 1.2 x article 2; flagellum 7-articulate, without callynophore. Antenna 2 peduncular article 3 0.5 x article 4; flagellum 7-articulate. Mandible incisors with slightly convex margins; palp article 2 slender , 1 x article 3, with 1 submarginal posterodistal A2-seta, D2-setae present , article 3 length 5.6 x breadth, with 1 distal D3-seta and 3 apical E3-setae. Maxilla 1 outer plate with setal-teeth in a 2/5 FIGURE 9. Sophrosyne integricauda sp. nov. , holotype female, 3.2 mm, MV J60739 , Cape Wickham, King Island, Bass Strait, Australia. Scales for U1–3 and T represent 0.01 mm, remainder represent 0.2 mm. arrangement . Maxilla 2 inner plate narrow, outer plate broader; inner plate length 0.6 x outer plate. Maxilliped inner plate subrectangular, without apical nodular robust setae; outer plate submarginal setae short, simple; palp article 2 slender, length 2.2 x breadth, 1.1 x article 3, article 3 slender, length 2.4 x breadth, dactylus with unguis absent. Gnathopod 1 basis length 1.5 x breadth, anterior margin with slender setae ; ischium length 1.2 x breadth; merus, posterior margin lined with long slender setae; carpus length 0.5 x propodus; propodus length 1.7 x breadth, posterior margin slightly concave, with robust setae, palm margin sinusoidal , posterodistal corner with produced subacute spine bearing vestigial seta . Gnathopod 2 minutely subchelate; ischium length 3.1 x breadth; carpus long, length 3.3 x breadth, posterior margin straight; propodus subrectangular, long, length 2.1 x breadth; palm transverse, with straight margin , with minute robust setae. Pereopod 3 merus weakly expanded anteriorly. Pereopod 4 merus weakly expanded anteriorly. Pereopod 5 basis moderately expanded posteriorly; merus moderately expanded posteriorly. Pereopod 6 coxa small, strongly lobate posteriorly ; merus slightly expanded posteriorly. Pereopod 7 basis posterior margin slightly rounded, minutely crenate, posteroventral margin rounded, posteroventral lobe not extending beyond ischium; merus not expanded posteriorly; propodus without setae along anterior margin. Epimeron 3 posterior margin with strong upturned spine above subquadrate posteroventral corner . Urosomite 1 with anterodorsal notch and rounded boss, without lateral flange. Uropod 1 rami subequal in length; outer ramus with 3 dorsal robust setae; inner ramus with 1 dorsal robust seta. Uropod 2 rami subequal in length; outer ramus with robust setae. Uropod 3 peduncle length 1.4 x breadth; rami subequal in length. Telson longer than broad, length 1.1 x breadth, moderately cleft (48%), distal margins incised, with 1 apical slender setae on each lobe. Remarks. Sophrosyne inverarae shares with S. murrayi and S. rodondo the same setal-tooth arrangement on the outer plate of maxilla 1, a vestigial seta on the posterodistal corner of the gnathopod 2 propodus, the shape of the basis of pereopods 5 and 6, a curved posteroventral corner with strong upturned tooth on epimeron 3 and a moderately cleft telson. Sophrosyne inverarae differs from both species in having D2- and D3-setae on the mandibular palp. It differs from S. murrayi most noticeably in having a minutely subchelate second gnathopod. Distribution. Loch Fyne, Inveraray, Scotland .