A review oI the Sphex flavipennis species group (Hymenoptera, Sphecidae) in northwest AIrica with description oI two new species and a discussion oI a species Irom Sardinia
Author
Schmid-Egger, Christian
christian@bembix.de
text
Linzer biologische Beiträge
2019
2019-07-26
51
1
459
471
journal article
22914
10.5281/zenodo.3758425
7ac1366d-0143-441f-89f5-4a0bbba2a48a
0253-116X
3758425
Sphex funerarius
GUSSAKOVSKIJ
,
1934
,
figs 3
(
4
,
10-13
)
Sphex funerarius
GUSSAKOVSKIJ, 1934: 3
, male, female. Lectotype: male
,
China: Gansu: Bei-lungshui (NRS)
.
M a t e r i a l e x a m i n e d:
5females
8 males
from
Tunisia
,
7 females
15 males
from Sardinia, around
100 specimens
from remaining Europe and palearctic Asia (all coll. CSE).
R e m a r k: The species occurs in the western palearctic region in three morpho- and colour
types
(for additional variation in central and eastern Palearctic region see MENKE & PULAWSKI 2000).
Females from Europe and western Asia (except Sardinia) have hindfemur black, red spots on femora are small, and red body colour is not orange but dark red. The females therefore resemble
S. maroccanus
. Males from this origin have apex of S8 with acute angle (figs. 10, 12, ca. 70 degree), the apex is somewhat pointed. Most specimens have
AS
5-10 with placoids, other have also a placoid on
AS
4. Number of placoids vary, see MENKE & PULAWSKI (2000) for details.
Females from Sardinia differ from the European mainland specimens as follows: apex of femora, tibiae and tarsi all red; red body colour is more orange red than dark red. These specimens are similar to the females from
Tunisia
. The apex of S8 of Sardinian males is larger than in other European males, and nearly right angled (fig. 11, 80-85 degree). Apical angle of S8 stands between
S. funerarius
males from European mainland and males of
S. maroccanus
. Males have placoids on
AS
5-9 (
9 specimens
) or on
AS
5-10 (
2 specimens
). Specimens from
Corsica
were not examined but probably also refer to this form, see MENKE & PULAWSKI (2000).
Females from
Tunisia
are similar in colour pattern to females from Sardinia (legs red apart most part of femora). S8 of these males looks similar to males from European mainland and palearctic Asian (fig. 13). Males have placoids on
AS
5-8 (
5 specimens
) or on
AS
5-9 (
2 specimens
).
Specimens from Sardinia and from European Mainland differ distinctly genetically by appr. 4.5% (fig. 1). This genetic gap in combination with the different form of male S8 gives a hint to a beginning species diversification of the Sardinian population. Because of this large genetic distance is a treatment as valid species of the Sardinian population also possible. However, the subject needs further examination in a larger geographic area, and especially the population from
Tunisia
has to be examined genetically, because they also may represent a different clade. The form from
Cyprus
is also distinctive and was described as
Sphex maxillosus
mavromoustakisi
de BEAUMONT, 1947. See MENKE & PULAWSKI (2000) for details.
D i s t r i b u t i o n:
S. funerarius
s.lat
is widespread in the whole western and central Palearctic region and reaches
Sweden
in the North. Recently the species is expansive in
Germany
, the northernmost location was
Berlin
in 2017 (pers. observation). Its distribution in North Africa is not clear. I could only examine
S. funerarius
from
Tunisia
, whereas all specimens from southern
Morocco
, formerly identified as
S. funerarius
, belong to
S. maroccanus
. It can be expected that
S. funerarius
is more widespread in northern Africa.