Review of the generaDiplopherusa and Paracladiscus (Coleoptera: Cleridae: Tillinae) from Japan and Taiwan, with descriptions of two new species Author Murakami, Hiroyuki text Acta Entomologica Musei Nationalis Pragae 2016 2016-11-15 56 2 725 741 journal article 10.5281/zenodo.5307482 0374-1036 5307482 D9281F86-E19D-4EA5-8173-DAB15ED8703B Paracladiscus choui sp. nov. ( Figs 6–7 , 15–16 , 22–24 , 29, 35, 41 , 51–52, 57–58, 61) Type materials. HOLOTYPE : J ‘[Taiwan] Dahanshan (Mt.) | Chunri Township | Pingtung County | [the locality name written in Chinese characters] | 8. VII, 2012 | H. Murakami leg.’ ( EUMJ ) . PARATYPES : TAIWAN : HSINCHU COUNTY: 5 JJ, Yulao , Alt. ca. 1,589 m , Jianshih Township , Hsinchu County , 3.vii.2015 , W. I. Chou leg. ( WCTT ) ; 3 JJ, same data, but the date is 4. vii. ( WCTT ) ; NANTOU COUNTY : 1 J, Songchuangang , Alt. ca. 2,500m , Renai Township , Nantou County , 25.vi.2014 , W. I Chou leg. ( WCTT ) ; PINGTUNG COUNTY : 1 J, same data as holotype ( TARI ) . TAITUNG COUNTY : 2 ♀♀ , Xiangyang , Alt. ca. 2,100 m , Haiduan Township , Taitung County , 1.vii.2012 , J. Yamasako leg. ( EUMJ ) ; 1 J, Yenping Forestroad , Yenping Township , Taitung County , 12.vii.2012 , H. Murakami leg. ( EUMJ ) ; 1 J, Dasyueshan , Alt. ca. 2,500m , Heping Township , Taichung County , 6.vii.2014 , by light trap with mercury lamp, S. Sin Yan leg. ( SSCT ) ; 1 J, Xiangyang , Alt. ca. 2,200 m , Haiduan Township , Taitung County , 25.vi.2015 , W. I. Chou leg. ( WCTT ) ; 1 J, Taimalishan , Alt. ca. 1,300 m , Taimali Township , Taichung County , 29.vi.2016 , W. I. Chou leg. ( WCTT ) . Description. Male ( Fig. 6 ). Body uniformly black, covered with brownish pubescence. Measurements (n = 3): EyW: 0.20–0.30 (0.25) mm; EyD: 0.50–0.62 (0.55) mm; BL: 4.45–5.91 (5.32) mm; PL: 1.10–1.47 (1.28) mm; PW: 0.71–1.01 (0.84) mm; EL: 3.35–4.43 (4.04) mm; EW: 1.00–1.32 (1.20) mm. Head closely and densely covered with setigerous punctures; frons elevated towards lateral margins; EyD/EyW: 2.00–2.50 (2.19). Terminal palpomeres of maxillary palpi short digitiform. Those of labial palpi longitudinal triangular, as long as width of apical margin of terminal palpomeres ( Fig. 22 ). Antennae ( Fig. 15 ) longer than total length of head and pronotum combined; antennomere I swollen and bent; II compact; III–X biramous; ramus becoming progressively longer towards V; XI flattened, fusiform. Approximate length ratio of antennomeres (n = 1): 3.3: 0.9: 3.4: 4.1: 4.2: 4.2: 4.2: 4.2: 4.2: 4.1: 6.7. Relative length ratio of rami of antennomeres III–X (n = 1): 1.0: 1.6: 1.8: 1.9: 1.8: 1.6: 1.4: 1.1. Pronotum widest before apex, constricted in basal 1/4; surface with large deep and densely arranged setigerous punctures; pronotal projections reaching to procoxal projection; PL/PW: 1.46–1.59 (1.54). Elytra elongate, subparallel-sided, widest at humeri, covered with large deep punctation; rows of punctures reaching to apex; distance among punctures slightly larger than puncture diameter; interspaces with pubescence; EL/EW: 3.35–3.42 (3.38); EL/PL: 3.02–3.42 (3.16); EW/PW: 1.32–1.59 (1.44). Metendosternite ( Fig. 23 ) with extended furcal arms anterolaterally; furcal laminae distally extended; process anteriorly extending from base of furcal arms. Unguis with basal denticles; pro and mesotibiae elongate, gently widened apically; metatibiae ( Fig. 24 ) elongate, gradually dilated towards basal half, thence slightly curved inwardly. Pygidium ( Fig. 29 ) rounded posteriorly; pygidial struts stout, anteriorly extended. Sternite VIII ( Fig. 35 ) weakly pigmented in medial portion; apical margin emarginate in middle. Spicular fork ( Fig. 41 ) with intraspicular plate. Aedeagus in fully extended condition (Figs 51–52) with finely spinose area between tegmen and phallus. Tegmen shorter than phallus; phallobase cylindrical without phallobasic struts, membranous in ventral view; phallobasic apodeme elongate, tapered posteriorly. Phallus divaricate in apical 1/5, dorsally with median orifice, and roundly pointed apex. Female ( Fig. 7 ). Measurements (n = 2): EyW: 0.20–0.22 mm ; EyD: 0.64–0.68 mm ; BL: 5.73–5.93 mm ; PL: 1.29–1.30 mm ; PW: 0.89–0.95 mm ; EL: 4.44–4.63 mm ; EW: 1.27–1.30 mm . Similar to male, but EyD/EyW: 3.09–3.20; antennae ( Fig. 16 ) crenate, their length equal to 2/3 of length of male antennae; hind tibiae simply elongate. Approximate length ratio of antennomeres (n = 1): 2.1: 1.0: 1.6: 1.8: 1.8: 1.9: 1.6: 1.5: 1.5: 1.4: 1.9. Relative width ratio of antennomeres (n = 1): 1.4: 1.0: 1.1: 1.4: 1.5: 1.8: 1.8: 1.8: 1.6: 1.6: 1.5. PL/ PW: 1.37–1.45; EL/EW: 3.49–3.56; EL/PL: 3.43–3.56; EW/PW: 1.37–1.43. Pygidium ( Fig. 57 ) almost straight at apical margin; pygidial struts stout. Sternite VIII ( Fig. 58 ) rounded posteriorly. Figs 55–60. Female pygidia (55, 57, 59) and 8th sternites (56, 58, 60). 55–56 – Diplopherusa shibatai ( Miyatake, 1965 ) ; 57–58 – Paracladiscus choui sp. nov. ; 59–60 – P. atricolor Miyatake, 1965 . Scale bars: 0.25 mm. Differential diagnosis. The new species can be distinguished from P. atricolor by antennomeres III–X biramous in males ( Fig. 15 ); females have antennae longer than combined lengths of head and pronotum; punctation of head and pronotum dense; elytral rows of punctures extending to apex; tibiae elongate in male. Etymology. The species name is dedicated to Wen-I Chou, who kindly helped and cooperated during a collecting trip to Taiwan in 2012. Distribution. Taiwan : Hsinchu Co., Nantou Co., Pingtung Co., and Taitung Co. ( Fig. 61 ).