A Revision Of The Indochinese Genera Cheirochela And Gestroiella Heteroptera Naucoridae And A Review Of The Tribe Cheirochelini Author Dan Author Polhemus Author John Author Polhemus Author Sites, Robert text Raffles Bulletin of Zoology 2008 2008-08-31 56 2 255 279 journal article 10.5281/zenodo.5340370 2345-7600 5340370 COPTOCATUS Montandon, 1909 Diagnosis. – Form elongate-oval. ( Fig. 1 ) Abdominal and thoracic venter with fine pile of short, closely appressed hairs. Pronotum not markedly expanded posteriorly (except in C . stereos ), posterior width less than 2× width of head; anterolateral angles of pronotum simple, not forming ventrallydirected cup-shaped depressions at apices. Hemelytra of brachypterous forms short and truncate, apices rounded, not reaching to abdominal tergite VI. Parameres variable in shape, either deeply cleft medially and asymmetrically bidentate, or uniramous and tapering on distal half, sometimes with a fringe of long setae on outer margins. Discussion. – The genus Coptocatus is confined to Borneo, and was proposed by Montandon (1909) to hold a single species, C . oblongulus , collected near Mt. Kinabalu. Five additional species have been described subsequently by Polhemus (1986) , Nieser & Chen (1991) and Zettel (2002) . Although clearly members of the tribe Cheirochelini based on synapomorphies of the head and ventral abdomen (see Polhemus, 1986 , Figs. 7, 8 ), Coptocatus species are narrower and more elongate in comparison to those of Gestroiella or Cheirochela (compare Figs. 1 , 2 and 18 ). The key below, modified from Zettel (2002) , will permit identification of the species described to date.