Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions Author Grego, Jozef 44C83AAB-CF4E-46E8-8146-DFBADA150AB6 Horná Mičiná, SK- 97401 Banská Bystrica, Slovakia. jozef.grego@gmail.com text European Journal of Taxonomy 2020 2020-07-27 691 1 56 journal article 10.5852/ejt.2020.691 918d86bb-f11d-414e-9c1e-21ec17737792 2118-9773 3961840 CC8487E0-169D-4B32-A2AB-F9B34DFB3F3B Plagigeyeria reischuetzorum sp. nov. urn:lsid:zoobank.org:act: A15894D4-DED0-4678-938A-FE39F32E0C51 Figs 8A , 9 G–J Type material Holotype BOSNIA AND HERCEGOVINA • 1 shell ; Republika Srpska, Bileća Municipality, Berkovići , Vrelo “Vrijeka” in Dabarsko Polje NW of Bjeljani ( Fig. 2 B–G); 43.07472222° N , 18.23916667° E ; 483 m a.s.l. ; Aug. 2009 ; Alexander Reischütz , Nicole Steiner-Reischütz and Peter L. Reischütz leg.; NHMW 111678 . Paratypes BOSNIA AND HERCEGOVINA • 49 shells ; same data as for holotype; NHMW 111679/1, HNHM-MOLL-104165/1 , coll. Reischütz /45, coll. Grego /2 • 21 shells ; same locality as holotype; Mar. 2010 ; Alexander Reischütz , Nicole Steiner-Reischütz and Peter L. Reischütz leg.; coll. Reischütz/19, coll. Grego/2 • 3 shells ; same locality as holotype; 31 Mar. 2018 ; J. Grego , D. Angyal , G. Jakaba and M. Olšavský leg.; SBMNH 626404 /1, coll. Grego /2 • 18 shells ; same locality as holotype; 22 Apr. 2019 ; J. Grego , G. Jakab , M. Olšavský and M. Kováčiková leg.; coll. Grego /16 . Other material BOSNIA AND HERCEGOVINA • 3 shells ; Republika Srpska , Bileća Municipality , Orahovice , Estavela “Obod” in Fatničko Polje; 22 Apr. 2019 ; J. Grego , G. Jakab , M. Olšavský and M. Kováčiková leg.; 43.018543° N , 18.348643° E ; 452 m a.s.l.; coll. Grego/3. Table 1. Shell morphometry comparison of Plagigeyeria lewarnei sp. nov. , P. mostarensis Kuščer, 1933 , P. pseudocostellina sp. nov. and P. reischuetzorum sp. nov.

Species	Specimen	H	W	BH	BW	AH	AW	CA	H/W	AH / AW	W /	H/	H/	W /	H/ ( W-BW)
BW	BH	AH	AW
mm	mm	mm	mm	mm	mm	°
Plagigeyeria	Holotype	2.79	1.52	1.66	1.10	1.51	1.24	43	1.84	1.22	1.38	1.68	1.85	1.23	6.64
mostarensis	from Kuščer 1933
Plagigeyeria	Hypotype	2.69	1.62	1.62	1.21	1.14	1.20	45	1.66	0.95	1.34	1.66	2.36	1.35	6.56
mostarensis	221236 SMF
Plagigeyeria	Topotype?	2.69	1.72	1.79	1.38	1.34	1.45	49	1.56	0.92	1.25	1.50	2.01	1.19	7.91
mostarensis	F1146 JG
Plagigeyeria	Buna	2.05	1.43	1.28	1.00	1.02	0.97	40	1.43	1.05	1.43	1.60	2.01	1,47	4.77
mostarensis var.	spring JG F0536
Plagigeyeria	HNHM	2.34	1.30	1.28	1.00	0.97	0.95	33	1.80	1.02	1.30	1.83	2.41	1.37	7.80
pseudocostellina	Holotype
sp. nov.	104164
Plagigeyeria	Paratype	2.10	1.55	1.34	1.03	1.00	0.86	32	1.35	1.16	1.50	1.57	2.10	1.80	4.04
pseudocostellina	JG F1140
sp. nov.
Plagigeyeria	Holotype	2.73	1.73	1.66	1.19	1.29	1.05	48	1.58	1.23	1.45	1.64	2.12	1.65	5.06
reischuetzorum	NHMW
sp. nov.	111678
Plagigeyeria	Paratype	2.51	1.73	1.66	1.19	1.36	1.19	47	1.45	1.14	1.45	1.51	1.85	1.45	4.65
reischuetzorum	HNHM
sp. nov.	104165
Plagigeyeria	Paratype	2.64	1.68	1.64	1.22	1.27	1.15	49	1.57	1.10	1.38	1.61	2.08	1.46	5.74
reischuetzorum	JG F1174
sp. nov.
Plagigeyeria	Paratype	2.42	1.69	1.53	1.19	1.16	1.16	47	1.43	1.00	1.42	1.58	2.09	1.46	4.84
reischuetzorum	JG F1173
sp. nov.
Plagigeyeria	Holotype	1.95	1.33	1.21	1.05	0.93	1.02	54	1.47	0.91	1.27	1.61	2.09	1.30	6.96
lewarnei sp. nov.	JG F0829

Dimensions Holotype : H 2.73 mm ; W 1.73 mm; BH 1.66 mm ; BW 1.19 mm ; AH 1.29 mm ; AW 1.05 mm . Paratypes : H 2.51 mm ; W 1.73 mm; BH 1.66 mm ; BW 1.19 mm ; AH 1.36 mm ; AW 1.19 mm . H 2.64 mm ; W 1.68 mm; BH 1.64 mm ; BW 1.22 mm ; AH 1.27 mm ; AW 1.15 mm . H 2.42 mm ; W 1.69 mm; BH 1.53 mm ; BW 1.19 mm ; AH 1.16 mm ; AW 1.16 mm . Etymology Named after the Austrian malacologists Alexander and Peter L. Reischütz with Nicole Steiner-Reischütz, who were the first to collect the species in the type locality. Description Shell is narrow-conical, 2.7 mm high with rounded apex and yellowish corneous periostracum, with five convex pagoduline whorls with rather weak suture. Shell surface finely axially ribbed at late whorls of the teleoconch. Faintly cancellate nepionic whorls covered by 25 close-set spiral ribs. Density of spiral ribs apically decreasing at protoconch to 15 and fading out at malleate nucleus. Space among the protoconch ribs is covered by characteristically granulated surface. Umbilicus narrow slit-like and obscured by the strongly reflexed columellar marginal fold. Larger expanding reflexed trumpet-like aperture irregularly oval and attached to body whorl by a parietal callus. Peristome sharp, and strongly outward reflexed along the aperture outline. Outer lip lateral profile and columellar lip profile characteristically sinuous. Expanded aperture conspicuously protruded against shell outline at basal view. Differentiating features The shells of P. reischuetzorum sp. nov. differ from the specimens of P. mostarensis known from the spring of the Bunica River near Hodbina ( Fig. 5 A–C) by their more inflated whorls and less sinuated labral and columellar margins, as well by a more prominent and a less declined aperture at the shell lateral outline. Shell morphometry comparison with the related species is presented in the Table 1 . Habitat The type specimens were found in sediments washed out from the cave spring “Sušica Pećina” and its side permanent spring Vrelo “Vrijeka” ( 483 m a.s.l.) situated 100 m eastward from the cave and rising from debris under the road to Fatnica. It is a part of northern Bregava Basin, likely draining the karst waters from Lukavačko Polje ( 895 m a.s.l.) into Dabarsko Polje ( 472 m a.s.l.). During the sampling in March 2018 the adjacent parts of Dabarsko Polje were flooded by large spring outlets creating a vast lake. Distribution The new species is so-far known only from the type locality.