Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)
Author
Páll-Gergely, Barna
Author
Hunyadi, András
0000-0003-3488-2817
hunand@freemail.hu
Author
Grego, Jozef
0000-0002-4977-0415
jozef.grego@gmail.com
Author
Reischütz, Alexander
0000-0002-8844-7004
alexander.reischuetz@gmx.at
Author
Auffenberg, Kurt
0000-0001-9660-6776
kurtauffenberg@gmail.com
text
Zootaxa
2021
2021-05-18
4973
1
1
61
journal article
6241
10.11646/zootaxa.4973.1.1
d0967b08-98d1-4ccd-b472-a678e0e57eda
1175-5326
4771958
42EB4BF2-A571-4894-9EEF-783649A27E4F
Dicharax verrucosus
Páll-Gergely & Hunyadi
n. sp.
Fig. 34
Material examined.
Holotype
:
Myanmar
,
Shan State
,
Taunggyi
, hill above
Aye Say Tee Pagoda
,
Dragon Cave
,
20°47.489’N
,
97°03.036’E
,
1380 m
a.s.l.
(locality code: 2018/44),
08 Oct. 2018
,
A. Hunyadi
,
K. Okubo
&
J.U. Otani
leg.,
HNHM 104862
(
holotype
: D:
2.83 mm
, H:
2.14 mm
)
.
Paratypes
:
10 shells, same data as holotype,
HA
.
Other material:
2 shells,
Myanmar
,
Shan State
,
Hsihseng
centre E ca.
6 km
, right side of rd. +
400 m
on unpaved rd., limestone hill,
20°7.983’N
,
97°18.145’E
,
1010 m
a.s.l.
(locality code: 2018/40),
07 Oct. 2018
,
A. Hunyadi
,
K. Okubo
&
J.U. Otani
leg.,
HA
.
FIGURE 34.
Dicharax verrucosus
Páll-Gergely & Hunyadi
n. sp.
, A: holotype (HNHM 104862), B: specimen from sample 2018/40 (coll. HA).
Etymology.
The specific epithet
verrucosus
(Latin for excrescence/wart) refers to the small swelling on R3, reminiscent of a wart.
Diagnosis.
A
Dicharax
species
with a relatively high spire, glossy, smooth R1, R2+R3 about a quarter whorl (R3 shorter than R2), and blunt swelling on R3.
Description.
Shell light grey, possibly translucent when fresh; shell outline rounded in dorsal view, spire elevated, body whorl rounded; protoconch low, rather glossy, 1.5 whorls; R1 approximately 2.25 whorls, smooth, glossy, with occasional, weak growth lines; boundary between R1 and R2 conspicuous due to dense R2 ribs; R2 ribs curved toward aperture, ca. 36–38 R2 ribs present; R2 ribs situated close to each other, spaces between ribs much less than rib width; R2+R3 covers ca. quarter whorl; R3 very short; about half as long as R2; sculpture of R3 similar to R1, boundary between R2 and R3 distinct, indicated by deep constriction; middle of R3 with prominent, blunt swelling; aperture slightly oblique to shell axis; rounded, with slight upper incision in parieto-palatal region; boundary between inner and outer peristomes distinct, especially at columellar portion; inner peristome slim, slightly protruding, outer peristome strongly thickened, expanded and reflected, especially toward umbilicus (in sample 2018/40 more conspicuously than in 2018/44); umbilicus relatively narrow, less than a third of shell width.
Measurements.
D:
2.76–3.24 mm
, H:
2.1–2.46 mm
(2018/44); D:
3.21–3.55 mm
, H:
2.46–2.5 mm
(2018/40).
Operculum.
Unknown.
Differential diagnosis.
The combination of the elevated spire, smooth R2, short R3 with a prominent swelling distinguishes this species from all congeners.
Distribution.
Both known populations occur in the southeastern part of
Shan
States,
Myanmar
. The straightline distance between the two is ca.
77 km
(
Fig. 17
).
Remarks.
The typical shells from Taunggyi are slightly smaller, and have less reflected peristomes than specimens from Hsihseng. However, since all other shell characters (shape, ratios of the shell regions, and shell sculpture) are identical, we treat them as the same species.