Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea) Author Páll-Gergely, Barna Author Hunyadi, András 0000-0003-3488-2817 hunand@freemail.hu Author Grego, Jozef 0000-0002-4977-0415 jozef.grego@gmail.com Author Reischütz, Alexander 0000-0002-8844-7004 alexander.reischuetz@gmx.at Author Auffenberg, Kurt 0000-0001-9660-6776 kurtauffenberg@gmail.com text Zootaxa 2021 2021-05-18 4973 1 1 61 journal article 6241 10.11646/zootaxa.4973.1.1 d0967b08-98d1-4ccd-b472-a678e0e57eda 1175-5326 4771958 42EB4BF2-A571-4894-9EEF-783649A27E4F Dicharax omissus (Godwin-Austen, 1914) Figs 29–30 Alycaeus omissus Godwin-Austen, 1914: 411 , Plate 155, Fig. 13 . Alycaeus ( Chamalycaeus ) omissus — Gude 1921: 231 . Material examined. Lectotype (hereby designated): 1 shell, “ Siam & Shan boundary, coll. Woodthorpe ”, NHMUK 1903.7 .1.1228.1 . Paralectotype : 1 shell, same data as for lectotype, NHMUK 1903.7 .1.1228. Other material: 1 shell ( Fig. 29B, D : 2.94 mm , H: 1.94 mm ) + 10 shells, Myanmar , Shan State , Hopong centre 5.8 km — Namsang , left side of rd. #4, near „Hopong Spring Cave”, 20°49.028’N , 97°13.469’E , 1110 m a.s.l. (locality code: 2018/35), 06 Oct. 2018 , A. Hunyadi , K. Okubo & J.U. Otani leg., HA ; 8 shells, Myanmar , Shan State , Taunggyi centre ca. 16 km — Hopong , rd. #4 N 1.5 km , „Shwe Pyi Aunchonda” monastery, 20°47.263’N , 97°8.239’E , 1110 m a.s.l. (locality code: 2018/46), 08 Oct. 2018 , A. Hunyadi , K. Okubo & J.U. Otani leg., HA ; 1 shell ( Fig. 30A, D : 2.81 mm , H: 1.8 mm ) + 7 shells, Thailand , Chiang Mai Province , 1.2 km N of Wat Tham Klaeb , 500 m side road from the junction, 19°33.918’N , 99°3.690’E , 650 m a.s.l. (locality code: 2015/08), 08 Feb. 2015 , A. Hunyadi leg., HA ; 1 shell ( Fig. 30B, D : 2.69 mm , H: 1.75 mm ) + 2 shells, Thailand , Mae Hong Son Province , 800 m from Tham Nam Lod toward the Buddhist temple, 19°33.928’N , 98°16.794’E , 650 m a.s.l. (locality code: 2015/12), 09 Feb. 2015 , A. Hunyadi leg., HA . FIGURE 29. Dicharax omissus (Godwin-Austen, 1914) . A: lectotype (NHMUK 1903.7.1.1228.1); B: site 2018/35. FIGURE 30. Dicharax omissus (Godwin-Austen, 1914) . A: site 2015/08 (coll. HA); B: site 2015/12 (coll. HA). Type locality. “ Siam and Shan boundary” . Diagnosis. A variable Dicharax species with a relatively densely, regularly ribbed R1, rarely with spiral striation, dense R2 ribs, a rounded central swelling on R3, and an expanded, but not reflected outer peristome. Measurements. D: 2.52–2.87 mm , H: 1.71–1.89 mm (2018/08 and 2015/12); D: 2.89–3.23 mm , H: 1.88–2.3 mm (2018/35 and 2015/46). Differential diagnosis. Typical Dicharax notus and D. omissus are very similar, and since only eroded specimens of the latter species were available, the distinction of these two species was somewhat questionable until newly collected material became available. These specimens reveal that D. notus is generally larger, R1 bears denser ribs (although some populations of D. notus have less dense R2 ribs), and its outer peristome is more reflected. Distribution. Dicharax omissus is known from the Shan States of Myanmar and neighbouring Thai provinces of Chiang Mai and Mae Hong Son ( Jirapatrasilp et al. 2021 ; Fig. 28 ). Remarks. The syntypes and the newly collected 4 samples (possibly 3 populations) show some variability in terms of the sculpture of R1 and R3, and the presence or absence of the notch at the basal portion of the aperture. However, all of these traits are rather minor and do not justify the distinction of species. The conchological differences of these populations are summarized in Table 3 . TABLE 3. Variation of shell characters of Dicharax omissus (Godwin-Austen, 1914) .

shell diameter (in mm)	R1 sculpture	R3 sculpture	aperture
typical omissus	?	without spiral striation	smooth	without basal notch
2015/08	2.61–2.87	without spiral striation	smooth	with basal notch
2015/12	2.52–2.84	without spiral striation	finely ribbed	with basal notch
2015/35, 2015/46	2.89–3.23	with spiral striation	finely ribbed	without basal notch

The shells of the two samples from Hopong have some slight spiral striation on the R1, which is highly unusual in Dicharax . The spiral striae of this species might not be homologous with those of other alycaeid genera, although this requires further investigation.