Provisional checklist of genera of the Pieridae (Lepidoptera: Papilionoidea) Author Braby, Michael F. text Zootaxa 2005 832 1 16 journal article 10.5281/zenodo.170669 cc432084-63d9-4ab2-a3aa-8e2efc7b740a 1175­5326 170669 Pierinae : Colotis group Klots (1933) tentatively divided the predominantly Afrotropical genus Colotis Hübner into four subgenera ( Colotis , Teracolus Swainson, Madais Swinhoe , Calopieris Aurivillius ), but noted that the genitalia among the first three showed no tangible characters. Ackery et al . (1995) subsequently discarded Teracolus , but treated Calopieris (not studied by Klots) as a distinct monotypic genus (with type species eulimene Klug ). Henning et al . (1997) created a new subgenus, Cuneacolotis G.A. Henning & S.F. Henning & Joannou & Woodhall, for Colotis agoye (Wallengren) but, since the authors noted that the taxon shares characters between Colotis and Teracolus , its status is highly questionable and it should be discarded. Stoneham (1957) divided the Afrotropical genera Eronia Hübner and Nepheronia Butler and proposed two new monotypic genera: Afrodryas Stoneham (with type species leda Boisduval ) and Lepteronia Stoneham (with type species pharis Boisduval ). These proposals have not been accepted by recent authorities (e.g. Larsen 1991 ; Ackery et al. 1995 ).