New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae)
Author
Brown, John W.
text
Zootaxa
2019
2019-09-17
4671
2
195
222
journal article
25519
10.11646/zootaxa.4671.2.2
978ec4b3-8d0e-4f47-8084-610b34b96759
1175-5326
3442344
8DA2FA3F-3629-4D10-92B0-671637D91DD4
Cybilla
Pogue
,
new genus
Type
species:
Phalonia hubbardana
Busck, 1907
, by monotypy.
Cybilla
Pogue
and Mickevich, 1990: 322 (nomen nudum).
Cochylini
New Genus 3 [
hubbardana
group]:
Brown 2005: 208
;
Metzler and Brown 2014: 278
.
Cybilla
is proposed for the single species
Phalonia hubbardana
, which traditionally has defied generic assignment. Described in
Phalonia
by
Busck (1907)
, it was treated as such by
McDunnough (1939)
and was placed in “incertae sedis” by
Powell (1983)
.
Pogue (1986)
proposed the name
Cybilla
, and
Brown (2005)
and
Metzler and Brown (2014)
treated it as “
Cochylini
New Genus”.
Pogue (1986)
concluded that
Cybilla
is most closely related to
Decuma
(now recognized as
Monoceratuncus
), and that the two are the most derived Cochylina genera in North America.
Diagnosis
.
Cybilla
shares with
Monoceratuncus
an unusually modified uncus, the absence of socii, and the absence of a median process of the transtilla.
Cybilla
can be distinguished from
Monoceratuncus
by the presence of small lateral plates fused to the base of the uncus, and by the 4-segmented maxillary palpi.
Description
. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus present; sensory setae of antenna ca. 1.0 times flagellomere diameter in male, shorter, sparser in female; labial palpus upturned, porrect, combined length of all segments ca. 1.2 times diameter of compound eye, segment III exposed; maxillary palpus 4-segmented. Thorax: Posterior crest absent; lateral scale tufts of metanotum flat. Forewing (
Figs. 5, 6
) length
4.6–5.6 mm
, length 3.0–3.1 times width, females usually slightly larger than males; costa straight, without costal fold in male, apex rounded; termen gently curved; Sc slightly less than 0.5 wing length; R
1
originating beyond middle of discal cell; R
2
originating nearer R
3
than R
1
; R
5
ending at costa; M
3
and CuA
1
separate; CuA
2
originating ca. 0.75 length of discal cell; CuP absent; A1+2 stalked at ca. 0.5 length. Hindwing moderately broad, costa straight, with costal roll extending about 0.4 length of costa in male; apex rounded; termen straight; Sc+R
1
ca. 0.65 wing length; Rs and M
1
stalked at 0.65–0.75 length of M
1
; M
3
and CuA
1
separate; CuA
2
originating ca. 0.65 length of discal cell; frenulum with one acanthus in male, two in female. Abdomen: Unmodified. Male genitalia (
Fig. 22
) with well-developed uncus bearing small, closely appressed, flat, lateral plates; socius absent; transtilla well developed, concave mesially, lacking median process; valva broad at base, attenuate distally, length ca. 2.1 times width; costa setose, apex produced, sacculus undulate from base to apex of valva, with patch of small spines at outer margin of first undulation from base; vinculum arms free distally. Phallus large, ca. 1.1 times length of valva, straight, parallel-sided in basal 0.33, enlarged in distal 0.5, with ventral spadelike projection; vesica with numerous cornuti. Female genitalia (
Fig. 36
) with papillae anales curved near middle, bulbous, slightly bilobed, setose; apophyses short, slender, length of apophyses anteriores ca. same as apophyses posteriores, ventral branch of apophyses anteriores joined to sterigma; sterigma a lateral band with prominent short-digitate, caudally-directed, subtriangular median process; ductus bursae slender, membranous throughout; corpus bursae small, rounded, with minute spicules; accessory bursa originating dorsomedially from corpus bursae.
Etymology
. The generic name is from the Latin goddess of nature, Cybele, and is feminine in gender.