Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae)
Author
Jałoszyński, Paweł
text
Zootaxa
2013
3750
5
549
568
journal article
10.11646/zootaxa.3750.5.7
d35e62c7-975b-4dff-892e-e8e5c926820e
1175-5326
217852
22C6D62A-D6CA-4BBE-AD0D-29174C2A2769
Alloraphes peruanus
Franz
(
Figs. 6
,
36–37
,
53–54
,
65
,
68
)
Alloraphes peruanus
Franz, 1980a: 214
, Fig. 200.
Type
material.
Holotype
:
Peru
(San Martín):
♂, three labels (
Fig. 65
): "Umg. Tarapoto /
Peru
, lg. Franz" with "SA133" on reverse side [white, printed; reverse handwritten in blue ink]; "
Alloraphes
/
peruanus
m. / det. H. Franz" [white, handwritten and printed]; "
Typus
" [red, handwritten] (NHMW).
Paratypes
(2 ♀; identity uncertain, see the Remarks section below): same data as
holotype
, except for yellow "
paratypus
" label (NHMW).
Revised diagnosis.
Male: frons and vertex confluent, without impression; pronotum with evenly arcuate antebasal transverse groove and without median pit; elytra unmodified; aedeagus in ventral view with nearly symmetrical apex of median lobe, long apical projections and narrow apices of parameres, one bearing two thin and short setae, the other one with a dense row of several thin and short setae. Female diagnostic characters unknown (see the Remarks section).
Redescription.
Body of male (
Fig. 6
) strongly convex, brown, covered with light brown vestiture; BL
0.83 mm
.
Head broadest at strikingly large, strongly convex and coarsely faceted eyes, HL
0.13 mm
, HW
0.20 mm
; tempora absent, eyes adjacent to occipital constriction; vertex and frons confluent, uniformly and weakly convex, anterior part of frons slightly flattened; supraantennal tubercles feebly marked. Punctures on frons and vertex inconspicuous, barely discernible; setae in the only known specimen almost entirely missing. Only rudiments of antennae preserved (left scape and right scape and pedicel).
Pronotum only slightly elongate, broadest in anterior fourth; PL
0.23 mm
, PW
0.21 mm
. Anterior margin and lateral margins in anterior third strongly rounded; sides distinctly but slightly narrowing caudad, pronotum distinctly constricted near posterior third; posterior margin slightly arcuate; ante-basal transverse groove distinct and evenly arcuate, without median pit and indistinctly deepened at each end. Punctures on pronotal disc inconspicuous, fine and sparse; setae on disc not preserved, on sides of pronotum visible remains of moderately long and disordered bristles.
Elytra more convex than pronotum, oval and unmodified, broadest slightly anterior to middle; EL
0.48 mm
, EW
0.38 mm
, EI 1.27; sub-humeral lines sharply marked and as long as 0.6 EL. Punctures on elytra distinct but small, shallow and with diffused margins, separated by spaces comparable to puncture diameters; setae on the only known specimen only partly preserved, relatively long, sparse and distinctly suberect.
Hind
wings well developed.
Legs long and slender; all tibiae straight.
Aedeagus (
Figs. 36–37
,
53–54
) slender and lightly sclerotized; AeL
0.25 mm
; in ventral view apical part of median lobe nearly symmetrical and with broad, shallow and asymmetrical emargination of apical margin; apical projections long; parameres in lateral view narrowing only in short and apical part which is curved dorsally, one bearing two thin and short setae, the other one with a dense row of several thin and short setae.
Female. Unknown (see the Remarks section).
Distribution.
Northern
Peru
(
Fig. 68
e).
Remarks.
The
type
series of
A. perunaus
is composed of a
holotype
male (partly damaged, with missing right elytron and most of the antennae) and three
paratypes
(one male and two females). The
paratype
male is not conspecific with
A. peruanus
and is fixed below as a
holotype
of a new species,
A. dentatus
sp. n.
Alloraphes peruanus
and
A. dentatus
are externally highly similar but differ clearly in several characters.
Alloraphes peruanus
has much larger eyes than
A. dentatus
, more uniformly convex vertex, which in
A. dentatus
is noticeably more convex in middle than on sides, and the pronotal ante-basal transverse groove is more evenly arcuate, while in
A. dentatus
it is slightly bi-arcuate with a median pit. In my opinion it is not possible to unambiguously assign the two remaining
paratype
females (one of them shown in
Fig. 7
) to either
A. peruanus
or
A. dentatus
. They both differ from males of
A. peruanus
and
A. dentatus
in distinctly broader elytra and smaller eyes; the characters associated with the vertex and pronotum are ambiguous. It seems more probable that the females are conspecific with
A. peruanus
and not with
A. dentatus
, based on additional collecting data given by Franz (1980a). The
holotype
and two
paratypes
were collected in Juan Guerra, in leaf litter of a relatively dry forest,
28.ix.1968
, while one
paratype
in Santana, in forest leaf litter,
27.ix.1968
. These two localities are in different parts of Tarapoto and the
paratype
from Santana is most likely the male described below as a new species.