The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits
Author
Huber, Bernhard A.
text
Zootaxa
2018
2018-03-19
4395
1
1
178
journal article
30485
10.11646/zootaxa.4395.1.1
8bee84d8-4a01-4e53-8eb3-af176ce1feb9
1175-5326
1202519
B43C234D-45C4-4A6D-9836-8A7524A5B291
Mesabolivar tapajos
sp. n.
Figs 11–12
,
58
,
61–62, 66–68
,
75–77
Diagnosis.
Males are easily distinguished from most known congeners (except
M. jamari
) by extremely long distal element on procursus (
Fig. 58
); from
M. jamari
by other details of tarsus and procursus (compare
Figs 57 and 58
; larger tarsal process, smaller widened part of procursus, more distal position of dorsal process of procursus); from
M. jamari
and other putatively close relatives also by armature of male chelicerae (
Figs 61–62
; cylindrical apophyses relatively slender), and by shape of epigynum (
Figs 66–67
,
75–76
; relatively small median depression with indistinct pocket; pair of conical lateral processes slender, tips slightly curved backwards).
Etymology.
The specific name is derived from the
type
locality; noun in apposition.
Type
material.
BRAZIL
:
Pará
:
♂
holotype
,
1♀
paratype
,
UFMG
(21505–06),
4♂
6♀
paratypes
,
ZFMK
(Ar 18957–58), Floresta Nacional de Tapajós, km 83 (
3.049°S
,
54.928°W
),
95 m
a.s.l.,
14–18.x.2016
(B.A. Huber, L.S. Carvalho).
Other material examined.
BRAZIL
:
Pará
:
1 juv.
, together with
paratypes
.
2♀
in pure ethanol,
ZFMK
(
Br
16- 258), same data as types
.
1♀
,
ZFMK
(
Ar
18959),
Floresta Nacional de Tapajós
, km 67, ‘site 2’ (
2.875°S
,
54.941°W
),
190 m
a.s.l.
,
15.x.2016
(
B.A. Huber
,
L.S. Carvalho
)
;
2♀
in pure ethanol,
ZFMK
(
Br
16-269), same data
.
1♂
3♀
,
ZFMK
(
Ar
18960),
Floresta Nacional de Tapajós
, km 72, ‘site 2’ (
2.940°S
,
54.933°W
),
140 m
a.s.l.
,
17.x.2016
(
B.A. Huber
,
L.S. Carvalho
).
Assigned
tentatively.
BRAZIL
:
Pará
:
1♀
,
ZFMK
(
Ar
18961),
Floresta Nacional de Tapajós
, km 221 (
4.046°S
,
54.938°W
),
80 m
a.s.l.
,
16.x.2016
(
B.A. Huber
,
L.S. Carvalho
).
Description. Male
(
holotype
)
MEASUREMENTS. Total body length 2.3, carapace width 1.1. Distance PME-PME 100 µm, diameter PME 110 µm, distance PME-ALE 100 µm, distance AME-AME 30 µm, diameter AME 40 µm. Sternum width/length: 0.75/ 0.55. Leg 1: 36.7 (8.6 + 0.5 + 8.8 + 16.9 + 1.9), tibia 2: 5.6, tibia 3: 3.9, tibia 4: 6.0; tibia 1 L/d: 93. Femora 1–4 width (at half length): 0.14, 0.14, 0.15, 0.14.
COLOR (in ethanol). Carapace ochre-yellow, with slightly darker lateral margins and large light brown median mark including posterior part of ocular area; clypeus not darkened; sternum medially slightly darker (orange to light brown); legs ochre, with darker rings on femora (subdistally) and tibiae (proximally and subdistally; indistinct), tips of femora and tibiae whitish; abdomen greenish-gray, with dark internal marks dorsally and laterally, small ventral mark behind gonopore.
BODY. Habitus as in
Fig. 11
; ocular area raised (higher than usual in the genus); carapace with distinct median furrow; sternum unmodified; clypeus swollen, whitish, with sclerotized rim.
CHELICERAE. With pair of long tapering apophyses with slender, slightly curved tips (
Figs 61–62
).
PALPS. In general very similar to
M. jamari
(cf.
Figs 55–56
), but slightly smaller (femur length: 0.64 vs. 0.72; femur maximum width: 0.26 vs. 0.32); only tarsus and procursus clearly different (
Fig. 58
): larger tarsal process, smaller widened part of procursus, more distal position of dorsal process of procursus.
LEGS. Without spines and curved hairs, few vertical hairs; retrolateral trichobothrium on tibia 1 at 2%; prolateral trichobothrium present on tibia 1; tarsus 1 with 25 pseudosegments, indistinct.
Male
(variation). Tibia
1 in
three other males: 8.9, 9.2, 9.7.
Female.
In general similar to male (
Fig. 12
) but dark mark on carapace larger, clypeus unmodified, dark rings on legs more distinct, sternum in most females with distinct light brown median mark. Tibia
1 in
nine females: 4.9– 5.6 (mean 5.2). Epigynum as in
Figs 66–67
,
75–76
; anterior plate with median depression bordered posteriorly by pocket; pair of conical lateral processes, tips slightly bent towards posterior; posterior plate simple. Internal genitalia as in
Figs 68
,
77
, with small pore-plates and distinctive median structure (curved median tube, possibly to accommodate long processes of procursi). One female from ~
110 km
S of
type
locality (“km 221”) with pocket in slightly more anterior position and shorter legs (tibia 1: 4.4); assigned tentatively.
Natural history.
The spiders were found close to the ground, in webs that were mostly hidden in the leaf litter, often in large rolled up leaves. Males and females were sometimes found together, hiding in the back of the shelter.
Distribution.
Known from Floresta Nacional de Tapajós in
Pará
state (
Brazil
) only (
Fig. 722
).
Note.
This species is possibly identical to “sp. 06” in
Machado (2011)
, described from a single male specimen from Altamira (~
300 km
E Tapajos).