Revisiting the taxonomy of the names Cephalotaxus mannii and C. griffithii (Taxaceae)
Author
Bisht, Sunita
University School of Environment Management, Guru Gobind Singh Indraprastha University, Sector 16 C, Dwarka,
Author
Khuraijam, Jibankumar Singh
0000-0003-4603-0216
Botanic Garden Division, CSIR-National Botanical Research Institute, Rana Pratap Marg, Lucknow- 226 001, India. jskhuraijam @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4603 - 0216
jskhuraijam@yahoo.com
Author
Singh, Rita
University School of Environment Management, Guru Gobind Singh Indraprastha University, Sector 16 C, Dwarka,
text
Phytotaxa
2021
2021-05-19
501
1
189
194
http://dx.doi.org/10.11646/phytotaxa.501.1.10
journal article
4427
10.11646/phytotaxa.501.1.10
f1e63d5e-979a-4ed6-b7b0-e4686a61a4b8
1179-3163
5424696
Cephalotaxus mannii
Hook.
f. (1886: t. 1523)
Lectotype
(
Step
I
.
Lang
et al.
(2013)
.
Step II
. designated here):—
INDIA
.
Meghalaya
: Khasi Hills, Lankhla Woods,
July 1885
,
G
. Mann s.n.
(
K000288019
, digital image!; isolectotype
CAL447046
!) (
Fig.1
).
Residual
syntypes
:—
INDIA
.
Meghalaya
:
Khasi Hills
,
Lankhla Woods
,
01 November 1885
,
G
.
Mann
s.n.
(
DD
!), (
CAL 447050
!), (
K000288007
,
K000287675
,
K001325156
, digital images!), (
A00003307
, digital image!), (
GH00003308
, digital image!), (
E00112582
, digital image!), (US00012030, digital image!), (
P00731282
, digital image!)
, (
PH
00004039, digital image!).
Distribution
:—
India
(
Meghalaya
,
Nagaland
,
Manipur
),
China
(
Guangdong
,
Guangxi
,
Yunnan
,
Tibet
), Northern
Myanmar
,
Laos
(
Khammouan
,
Vientiane
),
Thailand
,
Vietnam
.
Notes:—
The species is distinguished from
C. griffithii
by several distinctive morphological characters. Leaves in
C. mannii
are borne 70º–90º to the branchlets where as it is 30º–45º in case of
C. griffithii
.
Leaves of
C. mannii
are
1.5– 6 cm
long,
2–4 mm
wide and in
C. grifithii
leaves are
4–9 cm
long and
4–7 mm
wide. One of the main distinguishing character between the two species is the leaf base. Leaf base of
C. griffithii
is broadly rounded whereas
C. mannii
leaf have narrow base. Seeds of
C. griffithii
are
3.5–4.5 cm
long and in
C. mannii
seeds are
1.8–2.8 cm
long.
Second-step lectotypification:—
In the protologue of
C. mannii
,
the provenance was quoted as: “Khasia Mts., in Lankhla woods, about
5,000 ft.
,
Gustav Mann
”. During the study, we have traced out multiple sheets of (thirteen specimens) this homogenous collection (by Gustav Mann) housed at different herbaria.
Farjon (2010
,
2017
) and
Lang
et al.
(2013)
considered the specimen at Kew bearing “
1 November 1885
, G. Mann s.n.” as a
holotype
. Since there was not
one specimen
or illustration indicated or used by Hooker, there can be no
holotype
according to Art. 9.1 (
Turland
et al.
2018
). Nor can their use of “
holotype
” be corrected to “
lectotype
” under Art. 9.10, as they did not meet requirements of Art. 7.11 (Art. 9 Note 6), hence, their reference is therefore considered as effective first step lectotypification. So, the specimen selected here to be treated as second step
lectotype
.
FIGURE 1.
Lectotype of
C. mannii
,
K000288019! © The Board of Trustees of the Royal Botanic Gardens, Kew. Reproduced with the consent of the Royal Botanic Gardens, Kew.
FIGURE 2. A.
C. mannii
branch bearing immature green coloured pollen cones, Meghalaya;
B.
C. griffithii
foliage, Manipur (Inside picture: Pollen cones).
Notes on conservation status:—
Cephalotaxus mannii
is listed as Vulnerable in the IUCN Red List (
Liao & Yang 2013
). In the eastern Himalayan region of
India
, populations of
C. mannii
and
C. griffithii
are threatened by shifting cultivation and forest fire. In
Mizoram state
, the timber of
C. griffithii
is used for house building and making furniture (
Bhardwaj & Gakhar 2008
). However, in most of the other states in Northeastern
India
, populations of
C. griffithii
(
Fig. 2A
) remained unexplored due to remoteness of location and limited accessibility of the habitat of this species (
Moirangthem
et al.
2014
).