The Belisana spiders (Araneae: Pholcidae) from Tibet, China Author Zhu, Wenhui College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China & zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670 Author Yao, Zhiyuan College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China & yaozy @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 1631 - 0949 Author Zheng, Guo College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China & zhengguo @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 9735 - 9700 Author Li, Shuqiang Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China text Zootaxa 2020 2020-06-22 4802 1 111 128 journal article 10.11646/zootaxa.4802.1.7 1175-5326 3904926 46448DF5-3090-4A4C-9893-F6E68DB8636C Belisana gyirong Zhang, Zhu & Song, 2006 Figs 1–2 , 11 A–B Belisana gyirong Zhang, Zhu & Song 2006: 200 , figs 25–31. Material examined. 2 males ( IZCAS Ar39796–39797) and 2 females ( IZCAS Ar39798–39799), Chongse Village ( 28°23'42.02"N , 85°21'28.67"E , elevation 2801 m ), Gyirong Town , Gyirong County , Xigaze , Tibet , China , 25 July 2017 , X. Zhang and Z. Bai leg. Diagnosis. This species resembles B. zham sp. nov. ( Figs 9–10 , 11 G–H) with similar bulbal apophyses ( Fig. 2C ) and vulval pore plates ( Figs 2B , 11B ), but can be distinguished by presence of large retrolateral membranous flap (arrow f in Fig. 1D ), small, disto-dorsal membranous lamella (arrow 1 in Fig. 1C ) and small, disto-ventral membranous process (arrow 2 in Fig. 1C ) on procursus. Compared with B. zham sp. nov. , distal apophyses of male chelicerae (arrow da in Fig. 2D ) are closer, and epigynal pockets (arrow ep in Figs 2 A–B, 11A–B) are more separated. Redescription. Male (IZCAS Ar39796). Total length 2.35 (2.41 with clypeus), carapace 1.10 long, 0.83 wide, opisthosoma 1.25 long, 1.12 wide. Leg I: 21.73 (5.19 + 0.38 + 5.58 + 8.72 + 1.86), leg II: 14.40 (3.97 + 0.37 + 3.78 + 5.45 + 0.83), leg III: 9.87 (2.87 + 0.22 + 2.25 + 3.72 + 0.81), leg IV: 11.67 (3.59 + 0.26 + 2.82 + 4.04 + 0.96); tibia I L/d: 70. Habitus as in Figs 2 E–F. Carapace pale, with pair of large brown marks; ocular area and clypeus brown; sternum pale, without marks. Legs pale, without darker rings. Opisthosoma pale, with black spots dorsally and laterally. Distance PME-PME 0.10, diameter PME 0.07, distance PME-ALE 0.04, AME absent. Ocular area not elevated. Thoracic furrow absent. Clypeus unmodified. Sternum slightly wider than long (0.68/0.63). Chelicerae ( Fig. 2D ) with pair of small proximo-lateral apophyses and pair of short, curved distal apophyses each with hardened and pointed tip (distance between tips: 0.16). Pedipalps as in Figs 1 A–B; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with small proximo-dorsal apophysis (arrow in Fig. 1B ); procursus ( Figs 1 A–D) curved (arrow in Fig. 1A ), complex distally, with membranous elements, long distal spine and large, retrolateral membranous flap (arrow f in Fig. 1D ); bulb ( Fig. 2C ) with long hooked apophysis and long embolus. Retrolateral trichobothria of tibia I at 5% proximally; legs with short vertical setae on metatarsi, without spines or curved setae; tarsus I with 24 distinct pseudosegments. FIGURE 1. Belisana gyirong Zhang, Zhu & Song, 2006 , male. A–B. Pedipalp (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-dorsal membranous lamella, arrow 2 points at disto-ventral membranous process; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D). FIGURE 2. Belisana gyirong Zhang, Zhu & Song, 2006 , male (C–F) and female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H). Female (IZCAS Ar39798): Similar to male, habitus as in Figs 2 G–H. Total length 2.30 (2.44 with clypeus), carapace 0.89 long, 0.81 wide, opisthosoma 1.41 long, 0.97 wide. Tibia I: 3.78; tibia I L/d: 47. Distance PME-PME 0.11, diameter PME 0.06, distance PME-ALE 0.04, AME absent. Sternum slightly longer than wide (0.63/0.58). External female genitalia ( Figs 2A , 11A ) simple and flat, with pair of pockets 0.10 apart (ep in Figs 2 A–B, 11A–B). Vulva ( Figs 2B , 11B ) with slightly curved anterior arch and pair of elliptic pore plates. Variation: Tibia I in another male (IZCAS Ar39797): 4.49. Tibia I in another female (IZCAS Ar39799): 2.56. Natural history. The species was found on the underside of rocks. Distribution. China ( Tibet , Gyirong County, Gyirong Town. Fig. 12 ; the same as type locality see Zhang et al. 2006 ).