Typification of names in the genus Pinguicula L. (Lentibulariaceae)
Author
Domínguez, Yoannis
Universidade Estadual Paulista (Unesp), Faculdade de Ciências Agrárias e Veterinárias, Jaboticabal, Departamento de Biologia Aplicada à Agropecuária, Via de Acesso Prof. Paulo Donato Castellane s / n, CEP 14884 - 900, Jaboticabal, SP, Brazil.
Author
Valdés, Cristina Mercedes Panfet
Universidad de La Habana, Jardín Botánico Nacional de Cuba, Carretera del Rocío Km. 3, Calabazar, Boyeros CP 19230, La Habana, Cuba.
Author
Miranda, Vitor Fernandes Oliveira
Universidade Estadual Paulista (Unesp), Faculdade de Ciências Agrárias e Veterinárias, Jaboticabal, Departamento de Biologia Aplicada à Agropecuária, Via de Acesso Prof. Paulo Donato Castellane s / n, CEP 14884 - 900, Jaboticabal, SP, Brazil.
text
Phytotaxa
2017
2017-07-07
312
2
179
198
http://dx.doi.org/10.11646/phytotaxa.312.2.2
journal article
10.11646/phytotaxa.312.2.2
1179-3163
Pinguicula albida
Wright ex
Grisebach (1866: 162)
Protologue locality
:—
Cuba
occ.
Type
(
lectotype
, designated here):—
[
CUBA
].
Cuba
occ.[identali], 1865,
C
.
Wright
2885 [=561]
(
GOET
barcode
GOET004567
[digital photo!],
isolectotypes
US
No. 48177 barcode 00136344 [digital photo!],
WU
No. 075790 [digital photo!])
;
image of the
lectotype
is available at http://gwdu64.gwdg.de/pls/herbar/typen$typen.QueryViewByKey?
P
_
ID
=9199&
Z
_CHK=7133
).
Note
:—Charles Wright (1815–1885) collected plants in
Cuba
from 1856 to 1867 engaged by Asa Gray (
Howard 1988
). These collections have been widely discussed because many of the specimens, with a unique number, were derived from several gathering events; often from different localities or at different dates, in order to complete series of a single species. Hence, several examples of admixture material are known from Wright’s sheets. Asa Gray kept a series of Wright’s plants with the original labels and sent others to specialists for determination; afterwards he sold or distributed the remaining material to several herbaria (
Howard 1988
). Most of these materials have standard labels with the printed inscription “Plantae Cubenses Wrightianae, No., Coll.
C
. Wright” and sometimes with undeterminate localities (
Cuba
or
Cuba
oriental) and dates; however,
Pinguicula
specimens have no information on locality and just bear a collection year(s): 1865 or 1860–1864. Wright’s specimens related to new species described by
A
.
Grisebach (1860
,
1866
) sometimes bear two numbers assigned by Gray, a first one before shipping the material to Grisebach (e.g. 561 for
P. albida
); and the second, considered as the “Wright” number (species number, e.g. 2885 for
P. albida
), assigned after Grisebach’s determination. The “Wright” number was assigned to the distributed sheets of the “same taxon” disregarding locality and date of collection (
Rankin Rodríguez & Greuter 2000
). Moreover, the lack of specific information on locality and/or dates for many specimens complicates the typification process of Grisebach’s names based on Wright collections.
Howard (1988)
typified Grisebach’s new names selecting Wright’s specimens at
GOET
(Grisebach’s Herbarium) as “
holotypes
”. However, Howard’s typification appeared only on a microfiche appendix to his book, so it cannot be considered an effective publication according to Arts. 7.9 and 29.1 of the
ICN
(
McNeill
et al.
2012
). Several authors have dealt with Grisebach’s names typification after Howard’s typification (e.g.
Rankin Rodríguez & Greuter 2000
,
Graham 2005
,
Krings & Fantz 2006
,
Liede-Schumann & Meve 2008
), and all of them agree with Howard’s selection of
GOET
specimens as
lectotypes
, but the status of the remaining specimens distributed by Gray varies due to the doubts regarding whether all specimens with the same number were derived from a single gathering or not.
Rankin Rodríguez & Greuter (2000)
did not recognize any
isotype
for
Aristolochia linearifolia
Wright ex
Grisebach (1866: 115)
. On the other hand,
Graham (2005)
and
Liede-Schumann & Meve (2008)
considered to be
isotypes
all specimens with the same “Wright” number as the
holotype
(or
lectotype
), even though, each sheet may have been assembled with heterogeneous material. Regarding the typification of
P. albida
,
Casper (1966)
cited two Wright (2885) specimens as “
holotype
” (
P
) and “
isotype
” (
MO
) respectively. We have traced
10 specimens
of
P. albida
(
Wright 2885
)
at different herbaria (
G
,
GOET
,
HAC
,
MA
,
MO
,
NY
,
P
,
US
,
WU
and
YU
), and also we have confirmed the absence of Wright’s
Pinguicula
specimen at
GH
. However, other specimens could be found hereinafter in other collections. Hence, according to the Arts. 9.2, 9.3, 9.11 and 9.13 of the
ICN
(
McNeill
et al.
2012
) and following
Howard (1988)
, we herein designate as
lectotype
the specimen at
GOET
which bear a Grisebach handwritten label with the “original” number assigned by Gray (561). Moreover, among the remaining
nine specimens
, those bearing the same collection year (1865) as on the
lectotype
are considered by us as
isolectotypes
(Rec. 9
C
).