Revalidation of the stygobiotic species Haber zavreli (Hrabĕ, 1942) (Clitellata, Naididae, Tubificinae) with discussion on the closely related species Haber speciosus (Hrabĕ, 1931) and Haber monfalconensis (Hrabĕ, 1966) Author Dumnicka 1, Elzbieta Institute of Nature Conservation, Polish Academy of Sciences, al. Adama Mickiewicza 33, 31 - 120 Krakow, Poland Author Wojtal 1, Agata Z. Institute of Nature Conservation, Polish Academy of Sciences, al. Adama Mickiewicza 33, 31 - 120 Krakow, Poland text Subterranean Biology 2021 2021-09-13 39 143 156 http://dx.doi.org/10.3897/subtbiol.39.68397 journal article http://dx.doi.org/10.3897/subtbiol.39.68397 1314-2615-39-143 EE4A444AC6F941679D28156E5B45F963 9590FA7F138A5EE0995934DBFC644735 Haber zavreli ( Hrabe , 1942) Peloscolex zavreli Hrabe 1942: 23-26 (description of species, type locality: Rajec u Ziliny , in wells); Peloscolex speciosus (partim) ( Brinkhurst and Jamieson 1971 ): 514-515; Peloscolex zavreli ( Kasprzak 1973 ): 421-422 (short description of two specimens found in the wells in the Beskidy Mts, Poland) Haber zavreli ( Holmquist 1979 ): 52-53 (redescription); Peloscolex speciosus zavreli ( Hrabe 1981): 87-88 (distribution, short taxonomic discussion); Haber zavreli ( Brinkhurst 1981 ): 1062 (citation) Redescription. Almost all mature specimens without the posterior part of the body. Length of complete mature individuals: 10-11 mm, number of segments: 64-68. Body wall without papillae and without mucous cover, usually smooth, but in some specimens with tiny wrinkles (Fig. 1 ). Figures 1-3. Haber zavreli (Hrabĕ, 1942) 1 fragment of the body with hair seta and tiny wrinkles seen on dorsal side 2 shape of dorsal pectinate setae: 2A in V segment 2B in VII segment 2C in XV segment 3 shape of ventral setae: 3A in IV segment 3B in VIII segment 3C in XV segment. Singular long and smooth hair seta (Fig. 1 ) in all dorsal bundles (exceptionally two setae in anterior segments). Pectinate setae with long teeth in all segments (Fig. 2A-C ). The length of external and intermediate teeth is almost the same, but external ones are stouter. In anterior segments 5-7 intermediate teeth (Fig. 2A, B ) and distal end of the setae shovel-shaped, in posterior segments it is goblet-shaped with 3-4 intermediate teeth (Fig. 2C ). In the anterior dorsal bundles 1-2 setae, in the posterior segments - 1 seta. All ventral setae with the upper teeth longer than the lower (Fig. 3A-C ): in anterior segments about two times longer (Fig. 3A ), from segment VIII this difference is smaller (Fig. 3B, C ). The singular modified spermathecal and penial seta is present respectively in segments X and XI. They are thin and sharp-ended with hollowed distal part (Fig. 4A, B ) and they are inserted inside glandular sacs. The length of setae of mature individuals: hair setae up to 1000 µm long; dorsal anterior: 103-132 µm ; posterior: 87-103 µm ; ventral anterior: 230-243 µm ; ventral posterior: 148-160 µm ; spermathecal setae: 55-70 µm and penial setae 55-63 µm . Figure 4. Haber zavreli (Hrabĕ, 1942) A spermathecal seta in glandular sac B penial seta in glandular sac. Long, irregular sac-like ampullae of spermathecae (Fig. 5 ) sometimes reach IX segment, short spermathecal duct set off suddenly from ampulla and its ectal opening occurs slightly dorsally from the line of spermathecal setae. The long, "worm-like" spermatozeugmata either fill whole ampulla or are concentrated in its ental part. Male funnel small, vas deferens very long, coiled (Fig. 6a ) with distal part slightly thinner and shorter than proximal one. Proximal part of vas deferens narrower than tubular atrium (Fig. 7a ) and enters to it apically. Prostate gland small and compact enters to atrium almost medially (Fig. 7b ). Thin ejaculatory duct markedly sets off from the atrium (Fig. 7c ). Penial apparatus elongated with two, well visible bulges - one with basal membrane (the so-called "penis sheath"), and the second one with penial seta (Fig. 6b ). Figures 5-7. Haber zavreli (Hrabĕ, 1942) 5 Fragment of the body with spermathecal ampulla (marked by red frame) filled with long spermatozeugmata 6a fragment of vas deferens 6b penial apparatus 7 ectal part of male genital apparatus: a atria b prostate gland c ejaculatory duct.