Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species Author Motamedinia, Behnam Author Skevington, Jeffrey H. 0000-0002-1445-9870 jhskevington@gmail.com Author Kelso, Scott 0000-0001-5559-9477 scott.kelso@canada.ca text Zootaxa 2021 2021-07-16 5002 1 1 103 journal article 10.11646/zootaxa.5002.1.1 1175-5326 5122655 5DC2A66A-3F04-42D0-8A8C-F0686054E556 Tomosvaryella nodosa De Meyer, 1995 Figs 26A–D , 27I , 47C, 59 Diagnosis: This species can be recognized by a small projection in hind trochanter (Fig. 47C); epandrium wider than long in dorsal view ( Fig. 26A ); syntergosternite 8 slightly rounded in dorsal view ( Fig. 26A ); ejaculatory ducts coiled ( Fig. 26C–D ); gonopods equal in height ( Fig. 26B ); hypandrium slightly rounded, wider than gonopods in ventral view ( Fig. 26B ). Based on the shape of surstyli in dorsal and lateral view, this species resembles T. pruinosa Kozánek, 1992 . Tomosvaryella pruinosa lacks the small projection in hind trochanter. In addition, there is a gap between the surstyli and cerci of T. nodosa in lateral view ( Fig. 26C–D ). Meanwhile, It is similar to T. hamounensis sp. nov . See the diagnosis notes under T. hamounensis sp. nov . for more details. Specimens examined: IRAN : South Khorasan : Gholonabad , 33°10’N , 59°22’E , 1914 m a.s.l. , 11.v.2015 , B. Motamedinia , Malaise trap , JSS51962 ( 1♂ , CNC ) ; ISRAEL : Arava Valley , 0.2 km N. Hazeva Field School , small wadi, 30°46’N , 35°14’E , - 116 m a.s.l. , 30.iii.1995 , M. E. Irwin , Malaise trap , JSS52061 ( 1♂ , TAU ); Arava Valley , Iddan , wadi running east of date palm orchard, 30°48’N , 35°16’E , - 110 m a.s.l. , 16.iv.1995 , M. E. Irwin , Malaise trap , JSS52052 ( 1♂ , TAU ); 17.iii.1995 , M. E. Irwin , Malaise trap , JSS52054 ( 1♂ , TAU ); 24.iii.1995 , M. E. Irwin , Malaise trap , JSS51699 ( 1♂ , TAU ); 31.iii.1995 , M. E. Irwin , Malaise trap , JSS52053 ( 1♂ , TAU ); Arava Valley , Moshaav Hazeva , Wadi Shahak between agricultural fields, 30°46’N , 35°16’E , - 110 m a.s.l. , 12.iii.1995 , E. Irwin , Malaise trap , JSS51622 ( 1♂ , TAU ); 15.iii.1995 , M. E. Irwin , Malaise trap , JSS52060 ( 1♂ , TAU ); 19.iii.1995 , M. E Irwin , Malaise trap , JSS51730 ( 1♂ , TAU ); Arava Valley , Shizaf Nature Reserve , Hazeva , north of water treatment plant low sandy hummocks in small wadi, 30°46’N , 35°15’E , - 80 m a.s.l. , 16.iv.1995 , M. E. Irwin , JSS51698 ( 1♂ , TAU ); Arava Valley , small wadi, 2.4 km W. of hwy. 90 at km. 149, 30°42’N , 35°11’E , - 60 m a.s.l. , 16.iv.1995 , M. E. Irwin , hand net , JSS52058 ( 1♂ , TAU ); Arava Valley , Iddan Springs , in damp seep, east of spring, 30°49’N , 35°16’E , - 116 m a.s.l. , 21.iii.1995 , M. E. Irwin , JSS51811 ( 1♂ , TAU ); 31.iii.1995 , M. E. Irwin , JSS51812 ( 1♂ , TAU ); Haze- va Field School , 30°43’N , 35°15’E , 2.xi.1997 , S. Plotkin , Malaise trap , JSS52096 ( 1♂ , TAU ); Nahal Eshharim , 30°28’N , 34°35’E , 790 m a.s.l. , 2.v.1995 , I. Yarom , Malaise trap , JSS51790 ( 1♂ , TAU ); Nahal Mishmar , 31°24’N , 35°20’E , 11.iv.1994 , A. Freidberg & F. Kaplan , JSS51746 ( 1♂ , TAU ); B. S. Hazeva , 13.iii.1998 , S. Alfi , JSS51771 ( 1♂ , TAU ); Sniai Ofira , 27°52’N , 34°17’E , 22.iii.1981 ,A. Freidberg , JSS51721 ( 1♂ , TAU ); Taba , 29°30’N , 34°53’E , 14.v.2010 ,A. Freidberg , JSS51764 ( 1♂ , TAU ); Zin Wilderness Nakhal Zin at En Akrabim cane-covered sandy wadi, 30°53’N , 35°09’E , - 61 m a.s.l. , 7.iv.1995 , M. E. Irwin , Malaise trap , JSS51700 ( 1♂ , TAU ); UNITED ARAB EMIR- ATES : Abu Dhabi : Al Wathba Wetland Reserve , 24°15’N , 54°37’E , i.2015 , A. Saji & A. van Harten , Malaise trap , CNC470786 , CNC470791 , CNC470801 ( 3♂ , CNC ); ii.2015 , A. Saji & A. van Harten , Malaise trap , CNC470524 , CNC470585 , CNC470508 , CNC470473 , CNC470465 , CNC470545 , CNC470448–9 CNC470498 ( 4♂ , 5♀ , CNC ); iii.2015 , A. Saji & A. van Harten , Malaise trap , CNC470669 , CNC470688 , CNC470648 ( 2♂ , 1♀ , CNC ); ix.2013 , A. Saji & A. van Harten , Malaise trap , CNC470711–2 ( 2♂ , CNC ); al-Ajban, 24°22’N , 55°01’E , 5–12.vi.2006 , A. van Harten , Malaise trap , CNCD159561 ( 1♂ , USNM ); Al Bida’a Protected Area , 23°36’N , 53°58’E , 29.x.2018 , A. Saji & A. van Harten , Malaise trap , JSS52291–4 ( 4♂ , CNC ); Barqa Al Suqoor Protected Area , 23°60’N , 52°41’E , 29.iii.2018 , A. Saji & A. van Harten , Malaise trap , JSS52280–1 ( 1♂ , 1♀ , CNC ); 29.x.2018 ,A. Saji & A. van Harten , Malaise trap , JSS52283 ( 1♂ , USNM ); Wadi Shawkah , 25°04’N , 56°01’E , 5–12.v.2007 , A. van Harten , pan trap , JSS52125 ( 1♂ , EAD ) . FIGURE 26. Male genitalia of Tomosvaryella nodosa (CNC470508, CNC) A) dorsal view, B) ventral view, C, D) lateral view. Distribution: Egypt , Iran , Israel , United Arab Emirates ( Skevington 2020 ) ( Fig. 59 ). Note: Based on DNA barcodes, T. nodosa is similar to T. pruinosa , differing by 4.2%. (pairwise divergence) (Supplementary Table 1 ).