A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico
Author
Luna-Cozar, Jesús
jlcozar@yahoo.com.mx
Author
Anderson, Robert S.
jlcozar@yahoo.com.mx
Author
Jones, Robert W.
Author
León-Cortés, Jorge L.
text
Zootaxa
2014
2014-04-15
3788
1
1
63
journal article
5629
10.11646/zootaxa.3788.1
80d33caa-2379-4f5e-ba98-6d1cc29a7906
1175-5326
4913761
7484832A-025B-41FA-9696-DF7531C01AA4
26.
Tylodinus rufus
Luna-Cozar
,
sp. nov.
(
Figures 98
d-e, 76-77, 119, 134)
Diagnosis.
Length male,
2.8-3.3 mm
, female,
2.7-3.9 mm
. Width male,
1.3-1.5 mm
, female 1.4-2.0 mm. As for
T. complicatus
with following exceptions: body approximately 2x longer than wide, clothed only with yellow and narrowly elongate scales. Punctures on rostrum longer than punctures on head, apical area smooth. Pronotum red, in lateral view strongly convex. Elytra with basal margin slightly sinuate (
Figure 3
), tubercles on I2 and I3 red, declivity base with tubercle on I4 distinctly longer than tubercle on I2; striae 10 incomplete, surface with or without granules. Male genitalia (
Figure 98
d-e, 119) with median lobe moderately stout, apical process in lateral view abruptly curved downward. Female with sternite eight oblong, median membranous area as long as basal plate.
FIGURES 76–83.
Dorsal and lateral habitus:
76–77.
Tylodinus rufus
,
78–79.
Tylodinus dominicus
,
80–81.
Tylodinus noctis
,
82–83.
Tylodinus kuscheli
.
Comments.
The punctures on the pronotum are evenly distributed, but sometimes smooth and glabrous areas are present in the middle of the disc, which is occasionally weakly carinate. Male genitalia in
T. complicatus
,
T. rufus
,
T. noctis
and
T. dominicus
are very similar; however,
T. rufus
is recognizable from its allies by the pronotum coloration.
Geographic distribution.
México
(
Chiapas
).
Habitat and elevation.
Specimens were collected from leaf litter in high elevation cloud forest between
1900- 2950 m
.
Derivation of specific name.
Name derived from Latin
rufus
= red
Material examined.
Total
77 males
,
53 females
.
Holotype
male (
CMNC
):
México
:
Chiapas
,
San Cristóbal
de las
Casas
,
Cerro Huitepec
(pico) ca.
5 km
W San Cristobal
, elev.
2750 m
(
16° 44'33"N
,
92° 41'38.4"W
),
15.ix.1991
, coll.
R
.
Anderson
cloud forest litter, 91-101.
Paratypes
:
México
:
Chiapas
,
Coapilla
,
5 km
NE Coapilla
, elev.
1990 m
(
17.17602°N
93.13293°W
),
26.v.2008
, coll.
LLAMA
(Wm-A-04-1-01)
—
1 male
(
CMNC
); same locality (
17.17550°N
93.13227°W
),
LLAMA
(Wa-A-04-1-23)
—
1 male
,
1 female
(
CMNC
); 5
Km
NNW
Coapilla
, elev.
1915 m
(
17.18285°N
93.15112°W
),
24.v.2008
, coll.
LLAMA
(Wa-A-04-2-49)
—
1 female
(
CMNC
);
Cerro El Calvario
, near
Tapalapa
, elev.
2200 m
(
17° 11'11.8"N
,
93° 7'21"W
),
23.vii.2003
, coll.
R
.
Anderson
—
2 males
(
CMNC
); same locality, elev.
2000m
(
17° 10'25.7"N
,
93° 7'52.3"W
),
23.vii.2003
, coll.
R
.
Anderson
—
2 males
(
CMNC
);
Cerro de Tapalapa
, microwave towers, elev.
2260 m
(
17.18786°N
93.12308°W
),
27.v.2008
, coll.
R
.
S.
Anderson
—
1 female
(
CMNC
);
Coapilla
(6-8
Km
NE), elev.
2030 m
(
17° 10'25"N
,
93° 7'52"W
),
26.vii.2005
, coll.
R
.
Anderson
—
2 females
(
CMNC
);
Chalchihuitán
,
Cerros de Chalchihuitán
, elev.
2050 m
(
16° 59'20.8"N
,
92° 37'13.1"W
),
24.vii.2003
, coll.
R
.
Anderson
—
1 male
(
CMNC
);
Huixtán
,
Bazom
,
Barrio
los
Pinos
, elev.
2350 m
(
16.74689°N
92.48985°W
),
30.v.2008
, coll.
J. L. Cozar
—
2 females
(
ECOS
);
Merced Bazom
, elev.
2470 m
(
16° 44'39.5"N
,
92° 29'34.1"W
),
30.v.2008
, coll.
R
.
Anderson
—
1 female
(
CMNC
);
Tapalapa
,
Cerro de la Danta
, (
17° 11'14.3"N
,
93° 6'31.3"W
), 20.vii.94, coll.
R
.
Jones
—
1 female
(
ECOS
);
San
Juan Chamula,
Tzontehuitz
(
Pico
) ca.
10 km
NE San Cristóbal
, elev.
2950 m
(
16° 49'10.2"N
,
92° 35'3.8"W
),
16.ix.1991
,coll.
R
.
Anderson
—
2 males
(
CMNC
);
San Cristóbal
de las
Casas
,
15 km
E San Cristóbal
, elev.
2500 m
(
16.74708°N
92.48997°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-20)
—
1 male
,
1 female
(
CMNC
); same locality (
16.74689°N
92.48990°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-01)
—
4 males
,
3 females
(
CMNC
); same locality (
16.74687°N
92.48999°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-03)
—
1 male
(
ECOS
); same locality (
16.74686°N
92.49004°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-04)
—
1 male
(
CMNC
); same locality (
16.74703°N
92.49006°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-18)
—
2 males
,
1 female
(
ECOS
); same locality (
16.74701°N
92.49021°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-28)
—
1 male
(
CMNC
); same locality (
16.74698°N
92.49024°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-29)
—
1 male
(
CMNC
); same locality (
16.74690°N
92.49036°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05- 1-32)
—
1 male
(
ECOS
); same locality (
16.74695°N
92.49039°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-33)
—
1 male
(
ECOS
); same locality (
16.74707°N
92.490°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-1-38)
—
2 males
(
ECOS
);
Cerro Huitepec
(pico) ca.
5 km
W San Cristobal
, elev.
2750 m
(
16° 44'33"N
,
92° 41'38.4"W
),
15.ix.1991
, coll.
R
.
Anderson
—
1 female
(
CMNC
);
Cerro Huitepec
ca.
5 km
W San Cristóbal
, elev.
2700 m
(
16° 44'33"N
,
92°41'38.4"W
),
14.ix.1992
, coll.
R
.
Anderson
—
1 female
(
CMNC
);
Huitepec
, elev.
2328 m
(
16° 44'52.4"N
,
92° 40'23.2"W
),
17.vi.2004
, coll.
J. L. Cozar
—
1 female
(
ECOS
); same locality, elev.
2400 m
(
16° 44'12.5"N
,
92° 40'46.9"W
);
18.xi.1993
, coll.
R
.
Jones
—
1 male
,
1 female
(
ECOS
); same locality,
16.ii.1994
, coll.
R
.
Jones
—
3 males
(
ECOS
); same locality, elev.
2480 m
(
16.75176°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2- 02)
—
1 female
(
CMNC
); same locality (
16.75167°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-04)
—
1 male
(
ECOS
); same locality (
16.75145°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-09)
—
2 males
,
1 female
(
CMNC
); same locality (
16.75140°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-10)
—
2 males
(
ECOS
); same locality (
16.75127°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-13)
—
2 males
,
2 females
(
CMNC
,
CWOB
); same locality (
16.75122°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-14)
—
1 female
(
ECOS
); same locality (
16.75113°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-16)
—
1 male
(
CMNC
); same locality (
16.75109°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-17)
—
1 male
,
1 female
(
CWOB
); same locality (
16.75104°N
92.68270°W
),
29.v.2008
, coll.
LLAMA
(Wa-A-05-2-18)
—
1 male
(
CMNC
); same locality, elev.
2500 m
(
16° 44'42.7"N
,
92° 40'52"W
),
30.ix.1994
, coll.
R
.
Jones
—
2 males
—
1 female
(
ECOS
); same locality,
29.iv.1994
, coll.
R
.
Jones
—
1 male
(
ECOS
); same locality,
29.iv.1994
, coll.
R
.
Jones
—
1 male
(
ECOS
); same locality,
27.v.1994
, coll. B. Gómez-G y O. L. Gómez-N—
1 male
(
ECOS
); same locality,
27.v.1994
, coll. O. L. Gómez-N y B. Gómez-G—
1 male
(
ECOS
); same locality,
25.viii.1994
, coll. B. Gómez-G—
1 female
(
ECOS
); same locality,
24.i.1995
, coll.
R
.
Jones
—
1 male
(
ECOS
); same locality,
14.vi.1995
, coll. B. Gómez-G, M. Girón-I y A. Méndoza-R—
1 male
(
ECOS
);); same locality,
6.iii.1994
, coll.
R
.
Jones
—
1 female
(
ECOS
); same locality,
4.viii.1994
, coll. B. Gómez-G—
1 female
(
ECOS
); same locality, elev.
2668 m
(
16° 44'33"N
,
92° 41'19.3"W
),
9.vi.2004
, coll.
J. L. Cozar
—
1 male
(
ECOS
); same locality, elev.
2800 m
(
16° 44'35.2"N
,
92° 41'28.3"W
),
07.iv.1994
, coll.
R
.
Jones
—
1 female
(
ECOS
); same locality, elev.
2600 m
(
16.75163°N
92.68270°W
),
21.v.2008
, coll.
LLAMA
(Wa-A-05-2-05)
—
3 males
,
2 females
(
CMNC
,
CWOB
);
Huitepec, S
.
Cristóbal
, elev.
2480 m
(
16.75086°N
92.68270°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-2-22)
—
1 female
(
CMNC
); same locality (
16.75077°N
92.68270°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-2-24)
—
1 female
(
ECOS
); same locality (
16.75045°N
92.68270°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-2-31)
—
1 male
(
ECOS
); same locality (
16.75041°N
92.68270°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-2-32)
—
1 female
(
CMNC
); same locality (
16.75032°N
92.68270°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-2-34)
—
2 males
(
CMNC
); same locality (
16.75018°N
92.68317°W
),
28.v.2008
, coll.
LLAMA
(Wa-A-05-2-47)
—
1 female
(
CMNC
);
Reserva Huitepec
,
15.viii.1995
, coll.
R
.
Jones
—
1 male
(
UAQM
); same locality, elev.
2200 m
(
16° 45'50.4"N
,
92° 40'41.9"W
),
7.vii.2003
, coll.
R
.
Anderson
—
1 male
,
1 female
(
CMNC
); same locality, elev.
2300 m
(
16° 45'50.4"N
,
92° 40'41.9"W
),
15.vii.2005
, coll.
R
.
Anderson
—
1 male
,
1 female
(
BMNH
); same locality,
13.ix.2001
, coll.
R
.
Anderson
—
2 males
(
CMNC
); same locality, elev.
2400 m
(
16° 44'19"N
,
92° 29'18.2"W
),
11.vii.2007
, coll.
R
.
Anderson
—
5 males
,
5 females
(
BMNH
,
CMNC
,
ECOS
); same locality, elev.
2450 m
(
16° 44'19"N
,
92° 29'18.2"W
),
9.vii.2003
, coll.
R
.
Anderson
—
3 females
(
CMNC
); same locality,
11.vii.2007
, coll.
R
.
Anderson
—
1 female
(
CMNC
); same locality, elev.
2500 m
(
16°44'32"N
,
92°41'07" W
)
9.vi.2006
, coll.
U. Caballero
—
1 male
,
1 female
(
ECOS
); same locality,
17.vi.2006
, coll.
U. Caballero
—
1 male
,
4 females
(
ECOS
); same locality, elev.
2600 m
(
16° 45'0"N
,
92° 40'59.9"W
),
9.vii.2007
, coll.
M. G. Branstetter
—
6 males
,
2 females
(
CMNC
,
CWOB
); same locality,
9.vii.2007
, coll.
R
.
Anderson
—
2 males
(
CMNC
); same locality, elev.
2450 m
(16° 45'84"N, 92° 40’70"W),
11.vii.2003
, coll.
R
.
S.
Anderson
—
2 males
(
CMNC
); same locality, elev.
2750 m
(pico),
18.ix.1991
, coll.
R
.
S.
Anderson
—
1 male
(
CMNC
)
.