Taxonomic Revision And Morphology Of The Trapdoor Spider Genus Actinopus (Mygalomorphae: Actinopodidae) In Argentina Author Ríos-Tamayo, D. Author Goloboff, P. A. text Bulletin of the American Museum of Natural History 2018 2018-03-20 2018 419 1 84 journal article 0003-0090 Actinopus clavero , sp. nov. Figures 4A–H , 5A–D , 43 TYPE MATERIAL : HOLOTYPE ♂ (MACN-Ar 36532): Córdoba . Nono [ 31°46′ S 64°59′ W ], Jan. 1998 , P. Goloboff and C. Szumik , 1♂ . PARA- TYPES : 1♀ (MACN-Ar 36533), Córdoba . 15 km de Mina Clavero, Camino a Villa Carlos Paz [ 31°36′ S 64°54′ W ], Jan. 1981 , P. Grosso , A. Zanetic and P. Goloboff. Same data as the holotype, 1♂ (MACN-Ar 36534). FIGURE 4. Actinopus clavero , sp. nov. , male holotype (MACN-Ar 36532). A , carapace. B , palp. C–D , femur and trochanter III, spiniform bristles. E–G , copulatory bulb, three different views. H , sternum. Scale bars = 1 mm. ETYMOLOGY: The specific name is a noun in apposition that refers to the type locality. DIAGNOSIS: Females of A. clavero can be distinguished by the short and slender spermathecae (fig. 5C). Males can be distinguished by the slender and almost straight bulb (fig. 4E–G) with very subtle, almost invisible keels and apophyses; they also differ from A. reycali by the presence of prolateral spiniform bristles on trochanter and femora III (fig. 4C–D). DESCRIPTION: Holotype male (MACN-Ar 36532, figs. 4A–H). Total length: not taken because the abdomen is very compressed and deformed. Carapace (fig. 4A): length 3.72, width 3.24. Cephalic region: length 1.92, width 2.28; clypeus short, 10 bristles between each PME-PLE dyad; one larger medial bristle on both sides of the OQ. Fovea : length 0.66, width 1.44. Eyes, diameters and interdistances: AME 0.12, ALE 0.20, PME 0.12, PLE 0.08, OQ length 0.48, anterior width 1.84, 1.68 posterior width; AME-ALE 0.52, PME-PLE 0.24, AME-PME 0.28, ALE-PLE 0.24, AME-AME 0.28. Chelicerae: length 1.80, width 0.90, with internal dorsal bands of bristles reaching to the rastellum base. Cheliceral apex with 9 recumbent and elongated bristles that cover the base of the fang. Rastellum with 8 marginal blunt cusps and 3 small dorsal attenuated ones. Cheliceral furrow with 3 large promarginal teeth and 5 retromarginal and 5 denticles, near the promarginal teeth. Labium: length 0.60, width 0.66. Maxillae: anterior length 0.72; 1.08 posterior length, width 0.84. Sternum (fig. 4H): Center flat; length 2.58, maximum width 1.92, minimum width 0.66. Postlabial sigillum deep, posterior sigilla poorly defined and shallow. PMS: length 0.32, PLS: length of basal:medial:apical segments 0.40:0.28:0.20. Spigots : could not be observed. Lengths of legs and palp: I: 4.20, 1.62, 2.64, 3.00, 1.50, 12.96. II: 3.90, 1.62, 2.46, 2.88, 1.44, 12.30. III: 2.82, 1.62, 3.00, 3.12, 1.56, 13.20. Palp: 4.38, 2.28, 3.78, –, 1.02, 11.46. Chaetotaxy: Trochanter: III, 1 P SUP; I-II-IV, 0. Femora: I-IV, 0; III, 2-2/3-2-2 D. Patellae: I-II, 0; III, 11 P +9 on the margin, 1 P A, 2 D A +6 on the margin; IV, 36 P SUP-D ANT. Tibia: I, 1-1- 1-1 R INF, 2-1-1-2-2-2 V, 1-1-2 V ANT, 1-2-3-1- 1-2 V POST; II, 1-1-1-1-1 R INF, 2-1-1-2-2-2-V, 1-1-2 V ANT, 1-1-1-1-1-2 V POST; III, 1 D (1:2 B), with a crown of 19 thorns, 1-2-2 R, 1-1-2 V POST, 1-2 V ANT; IV, 2-2-2 v, 1-1-1 p, 1 r a. Metatarsi: I, 1-1-1-5-1 V, 1-1-1-2-1-3-3 V ANT, 1-1-1-2-5-2-2-3 V POST; II, 1-1-1-1-1-1 V, 1-1- 2-1-1-2 V ANT, 1-1-2-1-2-1-1-3 V POST; III, 1-1-2-1-1-2 P SUP-D ANT, 1-1-1-2 P, 1-2-2-2-1 V, 2-2-2-1-2 R; IV, 1-2-3 P, 1 D A, 1-1-2-2-4 R. Tarsi: I, 1-1-1-1 P INF, 2-1-1-1-1-4 R INF, 1-1-4 V; II, 1-1-1-1-1-1 P INF, 2-1-1-3-2-2-5 R INF, 1-1-1 V; III, 1-2-2-2 P INF (1:4 A), 2-2-2-2-1 R INF; IV, 1-3-2-2-3-2 P INF, 2 R A. Total number of retrolateral spines on tibiae I and II: 4 and 5 respectively. Palp: tibia long and slender (width/ length: 0.19±0.01) (fig. 4B). Bulb slim and slen- der, almost straight (fig. 4E–G). ATA barely visible; BTA little developed; slightly curved spermatic duct. Pseudoscopula: Tarsi: I–II, sparse (uniformly distributed throughout the segment); III–IV, dense and uniform. Metatarsi: IV, sparse 3:4 A. Trichobothria: Tibiae: I 4-6; II-III 4-5; IV 5-4. Metatarsi: I-II 9; III 10; IV 7. Tarsi: I 9; II 8; III-IV 6. Color: Carapace and chelicerae reddish brown. Booklungs with light markings. Sternum light brown, labium and maxillae darker. Legs like carapace. Spinnerets light yellow. FIGURE 5. Actinopus clavero , sp. nov. , female paratype (MACN-Ar 36533). A , carapace. B , tibia II, retrolateral. C , sternum. D , spermathecae. Scale bars = 1 mm. Paratype female (MACN-Ar 36533, figs. 5A–D): Total length: 17.28. Carapace (fig. 5A): length and width 5.40, 17 bristles on each posterior margin. Cephalic region: length 3.24, width 4.68; clypeus short with 25 bristles, 4 between AME-ALE, 13 bristles between PME and 9 behind each PME-PLE dyad; 7 on both sides of the OQ with one bigger central. Fovea: length 0.66, width 1.80. Eyes, diameters and interdistances: AME 0.12, ALE 0.32, PME 0.20, PLE 0.24, OQ 0.96, anterior width 3.00, 2.72 posterior width; AME-ALE 1.00, PME-PLE 0.20, AME-PME 0.48, ALE-PLE 0.44, AME- AME 0.28. Chelicerae: length 2.52, width 1.98, with bands of internal dorsal bristles reaching the rastellum base, dorsal anterior region with superficial grooves on 1:3 A. Cheliceral apex with 10 recumbent and elongated bristles covering the base of the fang. Rastellum with 12 marginal, blunt cusps, and 6 small dorsal ones. Fang groove with 6 large promarginal teeth and 5 retromarginal ones (a smaller one between 4th and 5th) and 8 denticles in the furrow, near the promargin. Labium: length 1.44, width 1.02, with 28 anterior cuspules. Maxillae: anterior length 1.38, 2.04 posterior length, width 1.38, width 1.40, with ~70 cuspules 1:3 V ANT, expanding to the anterior face. Sternum (fig. 5D): Center flat; length 3.78, maximum width 3.12, minimum width 1.14. Postlabial sigillum deep, posterior sigilla well defined. Abdomen: (poorly preserved) length 7.44, with small dark bristles. PMS: length 0.80; PLS: length of basal:medial:apical segments 0.88:0.40:0.40. Spigots: 3 large, 40 small on PMS; PLS, numbers of large(small) spigots on basal:medial:apical article 4(45):5(43):2(50). Lengths of legs and palp: I: 3.36, 2.10, 1.68, 1.92, 0.90, 9.96. II: 3.60, 2.10, 1.62, 2.10, 0.90, 9.32. III: 3.30, 2.40, 1.20, 2.40, 0.90, 9.90. IV: 4.20, 2.70, 2.10, 2.40, 0.90, 10.20. Palp: 3.00, 1.80, 1.92, –, 2.10, 8.82. Chaetotaxy: All femora: 0. Patellae: I-II, 0; III, 5 P SUP-D ANT, +11 on the margin, 2 R A, +5 on the margin; IV, 37 P SUP-D ANT +4 near the edge; palp, 1- 1 P. Tibiae : I, 1 P A, 1-1-1-1-1 R INF , 1-1 V POST ; II, 22 R INF, 1-1-1 V POST; III, 1 D B, with a crown of 19 thorns, 10 R; IV, 1 D A; palp, 8 P INF , 9 R INF . Metatarsi : I, 16 P INF , 13 R INF , 1-1-2 V POST ; II, 11 P INF, 12 R INF, 1-1-2 V POST; III, 2-1-3-1-1-2 D ANT, 19 on row D POST-R (more abundant R A); IV , 8 P (1:2 A), 1 D A ANT, 1-1 V ANT (1:3 A). Tarsi : I, 8 P INF , 12 R INF , 2 V A ; II, 11 P INF, 13 R INF, 1-2 V A; III, 13 R, 19 V-P; IV, 24 P-V; palp, 14 P INF , 18 R INF , 2 V A. Total number of retrolateral spines on tibiae I and II (fig. 5B): 5 and 22 respectively. Trichobothria : Tibiae : I-III 3-5 ; IV 3-4 . Metatarsi : I-II 9 ; III-IV 6. Tarsi: I 8; II 7 ; III-IV 5. Color : Carapace reddish brown, cephalic region and chelicerae darker. Abdomen light brown ; booklungs with light markings. Sternum light reddish brown, labium and maxillae darker. Spinnerets yellowish. Legs like carapace, tarsi dark . DISTRIBUTION: West of Córdoba province . ADDITIONAL MATERIAL EXAMINED: None.