A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia
Author
Zonstein, Sergei L.
urn:lsid:zoobank.org:author:EADD3607-30FF-49AE-93F5-8410630469BE
Steinhardt Museum of Natural History, Tel-Aviv University, 69978 Tel-Aviv, Israel
znn@tauex.tau.ac.il
text
European Journal of Taxonomy
2024
2024-10-24
967
1
185
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459
journal article
10.5852/ejt.2024.967.2699
2118-9773
13990819
C08B8027-50CC-417E-BCD4-5183B9FF6738
Raveniola insolita
sp. nov.
urn:lsid:zoobank.org:act:
A3E8F4A4-7067-4053-A329-41992E5E26D9
Figs 10
,
92
,
146
,
211
,
268
,
360
,
409–411
,
575
, 677, 752
Diagnosis
Habitually, as well as in the structure of the copulatory bulb, the only known male of
Raveniola insolita
sp. nov.
resembles males of
R
.
alajensis
sp. nov.
and
R
.
cucullata
sp. nov.
(
Figs 10
,
411
cf.
Figs 4–6
,
387
,
393–396
). However, it differs from the two latter species in having either a noticeably thinner proximal section of the embolus, additionally lacking a raised keel, or a considerably longer palpal tibia, compared with
R. alajensis
and
R. cucullata
, respectively (vs a thicker embolus provided with a long raised keel and a clearly shorter palpal tibia; see
Figs 360
,
409–410
cf.
Figs 352, 354–355
,
385–386
).
Etymology
The specific epithet
‘
insolita
’ is a Latin adjective (of the feminine gender) that means ‘odd, different, unusual’ and refers to the unusual male characters of the
holotype
(which are intermediate between those in
Raveniola alajensis
sp. nov.
and
R. cucullata
sp. nov.
, although the two latter species are not very closely related to one another).
Material examined
Holotype
TAJIKISTAN
•
♂
;
Peter I Mts
, environs of
Yashilkul
(also Yashnylkul)
Lake
; ca
39°07′ N
,
71°18′ E
;
3300–3400 m
a.s.l.
;
29 Jul. 1891
;
B. Grąbczewski
leg.;
ZISP
.
Description
Male
(
paratype
)
HABITUS
. See
Fig. 10.
MEASUREMENTS
. TBL 14.65, CL 6.58, CW 5.70, LL 0.49, LW 1.03, SL 3.38, SW 2.92.
COLOUR
. Carapace, palps and legs dark foxy brown (legs I–II slightly darker than palps and legs III– IV); eye tubercle blackish brown; chelicerae medium reddish brown; sternum, labium, maxillae and abdomen including spinnerets paler yellowish brown; darker chevron-like dorsal abdominal pattern medium brown.
CEPHALOTHORAX
. Carapace and chelicerae as shown in
Fig. 92
. Clypeus and eye group as in
Fig. 146
. Eye diameters and interdistances: AME 0.18(0.23), ALE 0.26, PLE 0.19, PME 0.18; AME–AME 0.16(0.11), ALE–AME 0.13(0.11), ALE–PLE 0.13, PLE–PME 0.05, PME–PME 0.42. Anterior cheliceral edge with 30–35 slightly thickened spikes. Each cheliceral furrow with 8 promarginal teeth and 3–4 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in
Fig. 211
. Maxillae with 30–31 cuspules each.
LEGS
. Tibia and metatarsus I as in
Fig. 268
. Scopula: entire distal on metatarsi I–II; entire on tarsi I–II; narrowly divided with setae on tarsus III; sparse and widely divided on tarsus IV. Trichobothria: 2 rows of 8–10 on tibiae, 14–18 on metatarsi, 15–16 on tarsi, 8–9 on cymbium. PTC I–IV with 9–12 teeth on each margin.
LEG
MEASUREMENTS
.
Femur
|
Patella
|
Tibia
|
Metatarsus
|
Tarsus
|
Total
|
Palp |
4.11 |
2.09 |
3.35 |
– |
1.01 |
10.56 |
Leg I |
6.09 |
3.15 |
4.28 |
5.11 |
2.78 |
21.41 |
Leg II |
5.56 |
2.93 |
4.12 |
4.43 |
2.79 |
19.83 |
Leg III |
4.82 |
2.47 |
3.02 |
4.35 |
2.57 |
17.23 |
Leg IV |
6.11 |
3.06 |
4.51 |
6.13 |
3.06 |
22.87 |
SPINATION
. Palp: femur d3, pd2; patella pd1; tibia d4, p3, r3, v6; cymbium d
~
25 spikes. Leg I: femur d4, pd3, rd3; patella p2; tibia p2, pv3, r3, rv2+2M; metatarsus v1. Leg II: femur d4, pd3, rd2; patella p3; tibia p2, v8; metatarsus p4(3), v6. Leg III: femur d4, pd3, rd2; patella p3, r1; tibia d2, pd3, p4, r3, v7; metatarsus d3, p4, r4, v7; tarsus p1(0), r1(0). Leg IV: femur d4, pd3, rd2; patella p2, r1; tibia d2, p3, r4(3), v9; metatarsus d2, p4, r4, v8; tarsus p1, r1. Tarsi I–II aspinose.
PALP
. Tibia, cymbium and copulatory bulb as shown in
Fig. 360
. Embolus with long basal portion provided with rudimentary keel and shorter hooked apical part (
Figs 409–411
).
SPINNERETS
. See
Fig. 575
. PMS: length 0.52, diameter 0.17. PLS: maximal diameter 0.56; length of basal, medial and apical segments 0.98, 0.60, 0.52; total length 2.10; apical segment triangular.
Female
Unknown.
Ecology
Judging from the label data, the
holotype
male was collected in an alpine meadow-steppe biotope at
3300–3400 m
a.s.l. (such as shown in
Fig. 677
). Other details remain unknown.
Distribution
Tajikistan
: the eastern part of Peter I Mts. See
Fig. 752
.