The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species Author Ho, - Z. text Zootaxa 2019 2019-10-18 4687 1 1 174 journal article 25190 10.11646/zootaxa.4687.1.1 de00ca6c-2843-4ae1-a39c-a29603947612 1175-5326 3508098 CE73264D-C234-4B82-A634-CAD6254C5957 Asymmetricata Ballantyne 2009 Figs 13 , 139−179 Asymmetricata Ballantyne in Ballantyne & Lambkin 2009: 30 (figs 5–8, 108–114). Yiu, 2012: 88 ; 2017: 88 . Fu 2014: 54 . Type species: Luciola circumdata ( Motschulsky 1854 ) designated by Ballantyne & Lambkin 2009: 30 . Diagnosis. A genus with relatively large species (usually 10 mm or longer) having a broad pronotum with laterally divergent margins, and convex sided elytra which are often brightly coloured, many with dark brown to black elytra having pale margins. Known larvae are terrestrial. Males distinguished from all other Luciolinae by the asymmetrical T8 which is emarginate on its left side ( Ballantyne & Lambkin 2009 fig. 7); all but one species have bipartite LO in V7. Females are macropterous and those examined have no bursa plates; in two species the median oviduct plate is subtriangular in outline; larvae have laterally explanate tergal margins ( Fu et al. 2012a: 22 ). Remarks . Asymmetricata was erected for two species transferred from Luciola ( As. circumdata Motsch. 1854 from Thailand and Cambodia , and As . ovalis [ Hope 1831 ] from India ) and characterised primarily by the asymmetrical male abdominal T8 ( Ballantyne & Lambkin 2009: 30 ). The species were keyed and distinguished by the nature of the light organs in V7 (either bipartite or entire). Two further species with a similar configuration of T8 and bipartite LOs in V7 viz. As . humeralis ( Walker 1858 ) comb. nov. and As . bicoloripes ( Pic 1927 ) comb. nov. are transferred from Luciola . Three species ( Luciola doriae Olivier 1885 , L . impressa Olivier 1910b and L. notatipennis Olivier 1909a ) are synonymised with As . humeralis comb. nov. However, the variability of the dorsal colour patterns indicated below could indicate further variability As . humeralis comb. nov. (interpretations were made from dead museum specimens where it may be difficult to interpret the original colour pattern). Assessing this variability requires examination of aedeagal and aedeagal sheath patterns. Ballantyne & Lambkin (2009) suggested the possibility of a gradation from entire to bipartite LO in V7 because of the anterior emargination of the LO in V 7 in As. circumdata . On the basis of specimens examined here it appears that the LO in V7 exists in two distinct forms either bipartite or entire (see below) but that there is no evidence of a gradation between the two different forms.