<document id="EDD7FD7F5B88B01DCBE446B36FAB7953" ID-DOI="10.11646/zootaxa.5005.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5141926" ID-ZooBank="1D4CEEED-12CD-4D80-BC43-16F7FFF23885" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627475353458" checkinUser="plazi" docAuthor="Quintero-Gutiérrez, Edwin Javier &amp; Halliday, Bruce" docDate="2021" docId="03C0C95AFFE4CE0DFF4E3028FD50F880" docLanguage="en" docName="zootaxa.5005.4.1.pdf" docOrigin="Zootaxa 5005 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neparholaspis pygmaeus Quintero-Gutiérrez &amp; Halliday 2021, new combination" docType="treatment" docVersion="2" lastPageNumber="427" masterDocId="FFF9B122FFFECE17FFD93606FFA0FFD9" masterDocTitle="Review of the mite family Parholaspididae Evans, 1956 (Acari: Mesostigmata)" masterLastPageNumber="459" masterPageNumber="401" pageNumber="427" updateTime="1699025819191" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="6463062EF21B6C321078B3506F436FFE">Review of the mite family Parholaspididae Evans, 1956 (Acari: Mesostigmata)</mods:title>
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<mods:namePart id="D4D734164C4A94BD1D6A60F20F11223E">Quintero-Gutiérrez, Edwin Javier</mods:namePart>
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<mods:date id="B36B54E25EC5ADAC26EFD22D88E9872A">2021</mods:date>
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<heading id="D09ECF20FFE4CE0DFF4E3028FC28F991" bold="true" box="[151,904,1582,1609]" fontSize="11" level="1" pageId="26" pageNumber="427" reason="1">
<emphasis id="B91DA45EFFE4CE0DFF4E3028FC28F991" bold="true" box="[151,904,1582,1609]" pageId="26" pageNumber="427">
96. 
<taxonomicName id="4C6903CFFFE4CE0DFF1E3028FD15F991" ID-CoL="9913f881-1054-4154-83b0-635a14265b9c" authority="(Ishikawa, 1980)" authorityName="Quintero-Gutiérrez &amp; Halliday" authorityYear="2021" baseAuthorityName="Ishikawa" baseAuthorityYear="1980" box="[199,693,1582,1609]" class="Arachnida" family="Parholaspididae" genus="Neparholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="427" phylum="Arthropoda" rank="species" species="pygmaeus" status="comb. nov.">
<emphasis id="B91DA45EFFE4CE0DFF1E3028FE48F990" bold="true" box="[199,488,1582,1609]" italics="true" pageId="26" pageNumber="427">Neparholaspis pygmaeus</emphasis>
(Ishikawa, 1980)
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<taxonomicNameLabel id="A22E1925FFE4CE0DFD623028FC28F991" box="[699,904,1582,1608]" pageId="26" pageNumber="427" rank="species">new combination</taxonomicNameLabel>
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<subSubSection id="C3732BC7FFE4CE0DFF4E3049FC21F95C" pageId="26" pageNumber="427" type="reference_group">
<paragraph id="8BD6784CFFE4CE0DFF4E3049FD27F9BE" blockId="26.[151,904,1582,1669]" box="[151,647,1615,1639]" pageId="26" pageNumber="427">
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<treatmentCitation id="0AC85E5DFFE4CE0DFF4E3049FD23F9BE" author="Ishikawa, K." box="[151,643,1615,1639]" page="162" pageId="26" pageNumber="427" year="1980">
<taxonomicName id="4C6903CFFFE4CE0DFF4E3049FD23F9BE" ID-CoL="8fc1a015-b787-4110-86fd-cff70ecd7caa" authority="Ishikawa, 1980 b: 162" authorityName="Ishikawa" authorityPageNumber="162" authorityYear="1980" box="[151,643,1615,1639]" class="Arachnida" family="Parholaspididae" genus="Gamasholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="427" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B91DA45EFFE4CE0DFF4E3049FE3FF9BF" box="[151,415,1615,1639]" italics="true" pageId="26" pageNumber="427">Gamasholaspis pygmaeus</emphasis>
<bibRefCitation id="EFF805BDFFE4CE0DFE7C3049FD23F9BE" author="Ishikawa, K." box="[421,643,1615,1639]" pageId="26" pageNumber="427" pagination="153 - 174" refId="ref29461" refString="Ishikawa, K. (1980 b) Taxonomic and ecological studies in the family Parholaspidae (Acari, Mesostigmata) from Japan (Part 3). Bulletin of the National Science Museum, Series A, Zoology, 6, 153 - 174." type="journal article" year="1980">Ishikawa, 1980b: 162</bibRefCitation>
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.
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</paragraph>
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<treatmentCitationGroup id="AB795F62FFE4CE0DFF4E306BFC21F95C" box="[151,897,1645,1669]" pageId="26" pageNumber="427">
<taxonomicName id="4C6903CFFFE4CE0DFF4E306BFE3FF95C" ID-CoL="8fc1a015-b787-4110-86fd-cff70ecd7caa" authorityName="Ishikawa" authorityYear="1980" box="[151,415,1645,1669]" class="Arachnida" family="Parholaspididae" genus="Gamasholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="427" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B91DA45EFFE4CE0DFF4E306BFE3FF95C" box="[151,415,1645,1669]" italics="true" pageId="26" pageNumber="427">Gamasholaspis pygmaeus</emphasis>
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.— 
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<bibRefCitation id="EFF805BDFFE4CE0DFE1C306BFD01F95C" author="Ishikawa, K." box="[453,673,1645,1669]" pageId="26" pageNumber="427" pagination="183 - 190" refId="ref29507" refString="Ishikawa, K. (1980 c) Taxonomic and ecological studies in the Family Parholaspidae (Acari, Mesostigmata) from Japan (Part 4). Reports of Research Matsuyama Shinonome Junior College, 11, 183 - 190." type="journal article" year="1980">
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, 1980c: 184
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; 
<treatmentCitation id="0AC85E5DFFE4CE0DFD75306BFCDDF95C" author="Ishikawa, K." box="[684,893,1645,1669]" page="91" pageId="26" pageNumber="427" year="1987">
<bibRefCitation id="EFF805BDFFE4CE0DFD75306BFCDDF95C" author="Ishikawa, K." box="[684,893,1645,1669]" pageId="26" pageNumber="427" pagination="91 - 93" refId="ref29622" refString="Ishikawa, K. (1987 b) The family Parholaspidae (Acarina, Meosostigmata) in Japan. In: Yang, H, Wang, Z., Jeffers, J. N. R. &amp; Ward, P. A. (Eds.), The Temperate Forest Ecosystem. ITE Symposium 20. Antu, Jilin province, pp. 91 - 93." type="book chapter" year="1987">
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, 1987b: 91
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.
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</paragraph>
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<materialsCitation id="3B017211FFE4CE0DFF4E30B2FACCF914" box="[151,1388,1715,1742]" country="Japan" location="Kuwabara" municipality="Matsuyama City" pageId="26" pageNumber="427" specimenCount="1" stateProvince="Ehime" typeStatus="holotype">
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locality and habitat:
</emphasis>
<collectingCountry id="F37E38DCFFE4CE0DFE0830B2FDB4F917" box="[465,532,1716,1742]" name="Japan" pageId="26" pageNumber="427">Japan</collectingCountry>
, 
<collectingRegion id="49ADB6AEFFE4CE0DFDC630B5FD44F914" box="[543,740,1715,1741]" country="Japan" name="Ehime" pageId="26" pageNumber="427">Ehime prefecture</collectingRegion>
, 
<collectingMunicipality id="6BB2E236FFE4CE0DFD3730B2FC09F914" box="[750,937,1715,1742]" pageId="26" pageNumber="427">Matsuyama City</collectingMunicipality>
, 
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, litter of 
<taxonomicName id="4C6903CFFFE4CE0DFB4930B2FACCF914" box="[1168,1388,1716,1742]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="26" pageNumber="427" phylum="Tracheophyta" rank="species" species="acutissima">
<emphasis id="B91DA45EFFE4CE0DFB4930B2FACCF914" box="[1168,1388,1716,1742]" italics="true" pageId="26" pageNumber="427">Quercus acutissima</emphasis>
</taxonomicName>
</materialsCitation>
. 
<materialsCitation id="3B017211FFE4CE0DFF4E30DEFE16F92B" box="[151,438,1751,1778]" collectionCode="NSMT" country="Japan" pageId="26" pageNumber="427" specimenCount="1" stateProvince="Ehime" typeStatus="holotype">
<emphasis id="B91DA45EFFE4CE0DFF4E30DEFEC0F928" bold="true" box="[151,352,1751,1778]" pageId="26" pageNumber="427">
<typeStatus id="54D2C6EEFFE4CE0DFF4E30DEFF72F92B" box="[151,210,1752,1778]" pageId="26" pageNumber="427">Type</typeStatus>
depository:
</emphasis>
<collectionCode id="ED78E089FFE4CE0DFEBE30DEFE16F92B" box="[359,438,1752,1778]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="26" pageNumber="427">NSMT</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD6784CFFE4CE0DFF4E311CFD50F880" blockId="26.[151,1437,1818,1881]" pageId="26" pageNumber="427">
<emphasis id="B91DA45EFFE4CE0DFF4E311CFF78F8ED" bold="true" box="[151,216,1818,1844]" pageId="26" pageNumber="427">Note:</emphasis>
We have transferred 
<taxonomicName id="4C6903CFFFE4CE0DFE1F311CFD49F8ED" authorityName="Ishikawa" authorityYear="1980" box="[454,745,1818,1844]" class="Arachnida" family="Parholaspididae" genus="Gamasholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="427" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B91DA45EFFE4CE0DFE1F311CFD49F8ED" box="[454,745,1818,1844]" italics="true" pageId="26" pageNumber="427">Gamasholaspis pygmaeus</emphasis>
</taxonomicName>
into 
<taxonomicName id="4C6903CFFFE4CE0DFCFB311CFC65F8ED" authorityName="Evans" authorityYear="1956" box="[802,965,1818,1844]" class="Arachnida" family="Parholaspididae" genus="Neparholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA45EFFE4CE0DFCFB311CFC65F8ED" box="[802,965,1818,1844]" italics="true" pageId="26" pageNumber="427">Neparholaspis</emphasis>
</taxonomicName>
because the peritrematal shields are fused to the ventrianal shield (not fused in 
<taxonomicName id="4C6903CFFFE4CE0DFDEA3138FD40F881" authorityName="Berlese" authorityYear="1904" box="[563,736,1854,1880]" class="Arachnida" family="Parholaspididae" genus="Gamasholaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="26" pageNumber="427" phylum="Arthropoda" rank="genus">
<emphasis id="B91DA45EFFE4CE0DFDEA3138FD40F881" box="[563,736,1854,1880]" italics="true" pageId="26" pageNumber="427">Gamasholaspis</emphasis>
</taxonomicName>
).
</paragraph>
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</treatment>
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