Review of Pennella Oken, 1816 (Copepoda: Pennellidae) with a description of Pennella benzi sp. nov., a parasite of Escolar, Lepidocybium flavobrunneum (Pisces) in the northwest Atlantic Ocean Author Hogans, W. E. text Zootaxa 2017 4244 1 1 38 journal article 36256 10.11646/zootaxa.4244.1.1 05f1a37b-29f3-4bca-a735-178a3373bb8a 1175-5326 400400 781D71C8-4632-4D1B-8D82-F77CA1146029 Pennella holteni ( Desmarest, 1825 ) Not valid. Desmarest (1825) named a new species while transferring the species described by Holten (1802) , P. exocoeti , from Lernaea to Lernaeopenna , both of which are synonyms of Pennella . There was no actual description of a new species in the original report. Accepted as P. exocoeti ( Walter & Boxshall 2015 ) .