Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa Author GUINOT, DANIÈLE Author TAVARES, MARCOS Author CASTRO, PETER text Zootaxa 2013 2013-05-29 3665 1 1 414 http://dx.doi.org/10.11646/zootaxa.3665.1.1 journal article 10.11646/zootaxa.3665.1.1 1175-5326 5276390 8358B363-BEE3-416D-96CA-8614E38B61D5 Family Corystidae In Corystes the male gonopore is coxal, with the short penis emerging from the wide gonopore on the slope of the inclined P5 coxa, just above its coxo-sternal articular condyle ( Fig. 8A ). The penis consists of a soft papilla, wide at its base and prolonged by a narrow tip, and lying very close to the broad lateral foramen of G1 endopodite. In the very narrow posterior part of the sternum, the two short, thick G1s are very close to each other, their wide bases covering the penis. Abdominal somites 1 and 2 are dorsal, and the G1 and G2 have a long protopodite, a condition related to the dorsal position of the first three abdominal somites. In C. cassivelaunus the G1 exhibits negative allometry, intensified in post-pubertal individuals ( Hartnoll 1972a: 149 , 153, fig. 6, table 5). The disposition is similar in the examined material of Gomeza and Jonas , the large and soft papilla emerging from a large gonopore (thus well protected) and lies very close to the broad lateral foramen of the G1 endopodite. See Monophyletic Heterotremata : Superfamily Corystoidea ).