A New Species of Forstercooperia (Perissodactyla: Paraceratheriidae) from Northern China with a Systematic Revision of Forstercooperiines Author Wang, Hai-Bing Author Bai, Bin Author Meng, Jin Author Wang, Yuan-Qing text American Museum Novitates 2018 2018-03-21 2018 3897 1 44 http://www.bioone.org/doi/10.1206/3897.1 journal article 4549 10.1206/3897.1 3e366419-f283-4f6b-aa2d-0b7618d37908 0003-0082 5373337 Forstercooperia ulanshirehensis , sp. nov. HOLOTYPE : A nearly complete cranium (AMNH 21608) ( figs. 2–4 ). REFERRED SPECIMENS: A pair of complete mandibles ( IVPP V20155 ) ( figs. 5 , 6 ), a fragmentary maxilla with P3–M3 ( IVPP V20154 ) ( figs. 7A, 7B ); a fragmentary mandible with the lower canine and p1–4 ( IVPP V20156 ) ( figs. 7C, 7D ), and a fragmentary mandible with p3–m3 ( AMNH 22101) ( fig. 8 ). TABLE 1. Measurements of cranium of Forstercooperia ulanshirehensis (mm). H = height; L = length; Max = maximum; W = width.

Cranium	AMNH 21608
Basilar L	420a
Profile L (= total length)	455 a
Occipital–postorbital process, L	236.0
Premaxilla–postorbital process, L	225 a
P1-condyle, L	376.0
Postglenoid process–condyle, L	96.5
Anterior end of orbital–nasal notch, L	117.3 a
Premaxilla–anterior end of orbital, L	185.5 a
Zygomatic, Max W	182 a
Palatine at P1, W	53.2 a
Palatine at M1, W	51.8 a
Palatine at M3, W	52.8 a
Postglenoid processes, Max W	131.5
Parocciptal processes, Max W	99.1
Occipital, H	120.5 a
Foramen magnum, Max W	24.9
Occipital condyles, Max W	76.3
Occipital condyle, H	42.9
Occipital condyle, W	35.9

a Approximate measurements. ETYMOLOGY: The specific name ulanshirehensis refers to the Ulan Shireh Formation, in which AMNH 21608 was collected. HORIZON AND LOCALITY: AMNH 21608 (field number, 613), Ulan Shireh Formation , Wulanhuxiu site (previously known as 8 miles north of Tukhum Lamasery ) of Erlian Basin , Nei Mongol , China ; AMNH 22101 (field number, 602), Ulan Shireh Formation , Wulantaolegai site (previously known as 4 miles north of Tukhum ) of Erlian Basin , Nei Mongol , China ; IVPP V20154 , V20155 , and V20156 , Irdin Manha Formation , Irdin Manha site, Erlian Basin , Nei Mongol , China . Middle Eocene. DIAGNOSIS: Small species of Forstercooperia , the length of upper premolar series about 81 mm , about 25% smaller than that of Forstercoopeia totadentata (~ 108 mm ), about 20% smaller than that of Forstercooperia mazhuangensis (~ 103 mm ) (see appendix, tables 2–3 ). Upper premolars transversely slightly narrower. Protocone and hypocone of P2 barely separate, metaloph of P2 narrow and posterolingually oriented. Metalophs (metaconules) of P3–4 short. Entoconid of p4 present.