Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan Author Shimizu, So Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan & DC and Overseas Challenge Program for Young Researchers, Japan Society for the Promotion of Science, Tokyo, Japan & Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK https://orcid.org/0000-0002-5202-4552 parasitoidwasp.sou@gmail.com Author Broad, Gavin R. Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK https://orcid.org/0000-0001-7223-5333 Author Maeto, Kaoru Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan text ZooKeys 2020 990 1 144 http://dx.doi.org/10.3897/zookeys.990.55542 journal article http://dx.doi.org/10.3897/zookeys.990.55542 1313-2970-990-1 7B73642C278D40F89091B26213C9A704 9F51F78CD53F5005A305DE65494002C4 Enicospilus dasychirae Cameron, 1905 Figure 15 Eniscospilus (sic) dasychirae Cameron, 1905a: 123; HT ♀ from Sri Lanka, NHMUK, examined. Eniscospilus (sic) horsfieldi Cameron, 1905a: 124; HT ♀ from Sri Lanka, NHMUK, examined; synonymised by Townes et al. (1961 : 273). Henicospilus borneensis Szepligeti , 1906: 138; HT ♀ from Borneo, TM, not examined; synonymised by Townes et al. (1961 : 273). Enicospilus nigrimarginalis Cushman, 1937: 311; HT ♂ from Taiwan, DEI, not examined; synonymised by Townes et al. (1961 : 274). Specimens examined. Total of 49 specimens (44♀♀5♂♂): Japan (8♀♀3♂♂), Sri Lanka (2♀♀), Taiwan (34♀♀2♂♂). Type series: HT ♀ of Eniscospilus (sic) dasychirae Cameron, 1905, Pundaluoya, SRI LANKA, I.1899, P. Cameron leg. (NHMUK, Type 3b.1267); HT ♀ of Eniscospilus (sic) horsfieldi Cameron, 1905, Peradeniya, SRI LANKA, IX.1902, P. Cameron leg. (NHMUK, Type 3b.1265). Distribution. Australasian, Eastern Palaearctic, and Oriental regions ( Yu et al. 2016 ); this is a predominantly Oriental species. JAPAN: [ Ryukyus ] Okinawa ( Chiu 1954 ; Gauld and Mitchell 1981 ; present study). Though this species is widely distributed in Southeast Asia, it is restricted to Okinawa in Japan. Bionomics. Recorded as a parasitoid of several species of Erebidae (subfamily Lymantriinae ) ( Gauld and Mitchell 1981 ; Chiu et al. 1984 ; Chen et al. 2009 ) and Noctuidae ( Tang 1990 ), although there are no host records from Japan. Differential diagnosis. This species is readily distinguishable from all other Enicospilus species by the unique small drop-shaped and isolated proximal sclerite of fore wing fenestra, as in Fig. 15F . Figure 15. Enicospilus dasychirae Cameron, 1905 ♀ from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing.