Fifty new genera of Hesperiidae (Lepidoptera) Author Cong, Qian Author Zhang, Jing Author Shen, Jinhui Author Grishin, Nick V. text Insecta Mundi 2019 2019-10-11 731 731 1 56 journal article 10.5281/zenodo.3677235 1ee19d5d-27cf-4c93-989c-3fe3eb467203 1942-1354 3677235 BA35690A-FC73-4E5A-A805-FE9550275FEC Livida Grishin , new genus http://zoobank.org/ 24179CB6-20EF-4DF2-B545-CFF25554444C Type species. Pythonides assecla Mabille, 1883 . Definition. A genus clearly near Gindanes Godman and Salvin, 1895 , Pythonides Hübner, [1819] , Quadrus Lindsey, 1925 , Zera Evans, 1953 and Ouleus Lindsey, 1925 , but without closer affinity to any one of them ( Fig. 5 ). The species included here in this genus were formerly placed in Pythonides , but are not monophyletic with the type species of this genus, Papilio jovianus Stoll, 1782 . Keys to E. 41.5 in Evans (1953) . Morphologically, distinguished from related genera by the combination of the shape of uncus (flattened at the tip with a short and thin central projection) with the shape of valva (broad at the base with concave costa and diamond-shaped harpe longer than the rest of the valva, with serrated dorsoposterior edge), only one upper spur on hind tibiae, the lack of tuft in hind tibiae and thoracic pouch in males, ventral hindwing mostly pale blue, dorsal hindwing pale blue in males at least near tornus. In DNA, a combination of the following base pairs is diagnostic: aly 1672.3.1 :T128G, aly 1672.3.1 :C110G, aly 2618.5.1 :C3626G, aly140.1.2:T1822T (not A), aly 1349.7.9 :C1010C (not G), aly1877.13.1:A785A (not C), aly23605.1.46:T5641T (not A), aly23605.1.46:C5642C (not G), aly536.102.1:C56C (not A), aly 1935.2.1 :G242G (not C), aly345.3.1:C328C (not G), aly767.12.13:C118C (not A). Etymology. The name is a feminine noun in the nominative singular, from the Latin lividus (blue), to indicate the distinctive cornflower blue color of the hindwing in males of the type species. Figure 5. Genomic trees of Achlyodini . See Fig. 2 legend for notations. Species included. The type species and Pythonides grandis Mabille, 1878 . Parent taxon. Subtribe Pythonidina Grishin, 2019 . Comments. Another genus that would be very difficult to detect without genomic data, because by appearance the adults of this species do not look noticeably different from Pythonides . However, genomic analysis suggests that the two are distant.