Review of the species of the genus Serrapinnus Malabarba, 1998 (Teleostei: Characidae: Cheirodontinae) from the rio Tocantins-Araguaia basin, with description of three new species
Author
Malabarba, Luiz R.
Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500. 91501 - 970 Porto Alegre, RS, Brazil.
Author
Jerep, Fernando C.
Universidade Estadual de Londrina, Programa de Pós-Graduação em Ciências Biológicas, Departamento de Biologia Animal e Vegetal, Centro de Ciências Biológicas, 86057 - 970 Londrina, PR, Brazil. fjerep @ gmail. com & Research Associate, Division of Fishes, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, D. C.
text
Zootaxa
2014
2014-08-06
3847
1
57
79
journal article
5344
10.11646/zootaxa.3847.1.3
17adfe62-71be-4ddf-bbe2-9137253a2007
1175-5326
4928753
93242732-2B5A-4083-8EC5-6FA53BD83E7D
Key to the species of the
Cheirodontinae
occurring in the Atlantic drainages of northeastern
Brazil
, from the rio Tocantins-Araguaia to the rio Paraguaçú drainages
1. Lower lobe of caudal fin of males covered with cone shaped or papilla-like organs (see
Malabarba
et al.
, 2004
: figs. 1 to 4). A conspicuous small black spot present at midlength of first branched anal-fin ray of males (see
Malabarba
et al.
, 2004
: figs. 5–6 and 8). A dark brown nearly isosceles triangular shaped mark present in the area covering pelvic bone in males (see
Malabarba
et al.
, 2004
: fig. 9)....................................................................2 (
Kolpotocheirodon
)
1´. Lower lobe of caudal fin of males not covered with cone shaped or papilla-like organs. Conspicuous black spots lacking on branched anal-fin rays. Triangular mark in area covering pelvic bone absent....................................... 3
2. Dorsal fin lacking conspicuous black spots. 3–5 very small, vertical bars crossing lateral body stripe between pseudotympanum and area ventral to dorsal fin (see
Malabarba
et al.
, 2004
: fig. 8). Twelfth to the 14
th
or 15
th
principal caudal-fin rays of males bearing hooks...................................................................................................
Kolpotocheirodon theloura
(headwaters of São Francisco and
Paraná
river basins near Brasília, DF,
Brazil
)
2´. Conspicuous small black spot present on soft tissue between midlength of first and second, and second and third dorsal-fin branched rays (see
Malabarba
et al.
, 2004
: figs. 5 and 6). Vertical bars crossing lateral body stripe and hooks on caudal fin absent.................................................
Kolpotocheirodon figueiredoi
(Paraguaçu River drainage)
3. Base of lower caudal-fin lobe with hypertrophied group of scales forming small pouch................................
Compsura heterura
(São Francisco River basin and northeastern coastal drainages; absent in Tocantins-Araguaia River basin)
3´. Base of lower caudal-fin lobe without hypertrophied scales................................................... 4
4. Ventral procurrent caudal-fin rays not hypertrophied or exposed along ventral margin of caudal peduncle of mature males. Thirteenth to 14
th
caudal-fin rays bearing hooks associated with hypertrophied skin flaps in mature males (
Malabarba & Weitzman, 1999
: figs. 3–6)....................
Acinocheirodon melanogramma
(São Francisco and Jequitinhonha river basins)
4´. Ventral procurrent caudal-fin rays hypertrophied and exposed ventrally in mature males. Caudal-fin rays without hooks or hypertrophied skin flaps in mature males.................................................................. 5
5. First through fourth or fifth branched anal-fin rays of mature males slab-shaped (
Malabarba & Jerep, 2012
: fig. 3). Distal tip of ventral procurrent caudal-fin rays conical in mature males (
Malabarba & Jerep, 2012
: fig. 3). Anal-fin rays iv–v, 14–17. Lateral line completely pored with 32–36 scales...................
Ctenocheirodon pristis
(Tocantins-Araguaia River basin)
5´. First through sixth or eighth branched anal-fin rays of mature males slab-shaped (
Fig. 4
). Distal tip of ventral procurrent caudal-fin rays spatulate or scimitar-shaped in mature males (
Fig. 2
). Anal-fin rays iii–iv,16–21.............. 6 (
Serrapinnus
)
6. Three central cusps of dentary teeth expanded relative to other cusps and form a sharp-cutting edge. Lateral line completely pored....................
Serrapinnus heterodon
(São Francisco River basin and northeastern Atlantic coastal drainages)
6´. Three central cusps of dentary teeth similar in shape to remaining cusps (
Figs. 3
,
7
,
9
,
11
). Lateral line incomplete, with 5–10 pored scales (variable in
S. tocantinensis
, ranging from 6–36 scales)............................................. 7
7. Longitudinal black stripe wide and conspicuous, one or one and half scale deep, extending from posterior margin of opercle to caudal spot (
Fig. 8
)..................................................
S. sterbai
(Tocantins-Araguaia River basin)
7´. Longitudinal black stripe narrow, extending from region below dorsal-fin base to caudal spot......................... 8
8. Longest unbranched dorsal- and pelvic-fin rays of males prolonged as filament (
Fig. 10
)...........................................................................................
S. tocantinensis
(Tocantins-Araguaia River basin).
8´. Longest unbranched dorsal- and pelvic-fin rays of males not prolonged as filament.................................. 9
9. Ventral procurrent caudal-fin rays 17–19................................
S. lucindai
(Tocantins-Araguaia River basin)
9´. Ventral procurrent caudal-fin 16 or fewer.................................................................. 10
10. Ventral procurrent caudal-fin rays of mature males spatulate, forming keel along posterior ventral margin of caudal peduncle......................................
S. piaba
(São Francisco River basin and northeastern coastal Atlantic drainages)
10´. Ventral procurrent caudal-fin rays of mature males scimitar-shaped and forming semicircle, with portion of ventral procurrent caudal-fin rays having resultant star-shape form with multiple points (
Fig. 2a
).................................... 11
11. Premaxillary teeth with 7 to 9 cusps (
Fig. 3a
)...............................
S. aster
(Tocantins-Araguaia River basin).
11´. Premaxillary teeth with 5 cusps (see
Jerep & Malabarba, 2014
: fig. 3)..........................................................................................
S. potiguar
(
Ceará-Mirim River
basin in
Rio Grande do Norte state
)
.