Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini)
Author
Malm, Tobias
Author
Johanson, Kjell A.
text
Zoological Journal of the Linnean Society
2008
2008-07-31
153
3
425
452
https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00403.x
journal article
10.1111/j.1096-3642.2008.00403.x
0024-4082
5446337
GRACILIPSODES
SYKORA, 1967
Gracilipsodes
Sykora, 1967: 594
;
Morse, 1981: 262
;
St Clair, 1994: 176
;
Ward, 2001: 73–82
.
Type
species:
Gracilipsodes psocopterus
Sykora, 1967
, by original designation.
Revised generic description, males
Adult:
Body and wing colour yellowish brown to dark greyish brown (denuded in alcohol).
Head:
Antennae with broadly oval scape, about as long as eye diameter (
Fig. 1A
). Frontal setal warts (f.w.) oval (
Fig. 1B
); anterior setal wart (a.w.) narrowly triangular or diamond shaped (
Fig. 1B, C
); anterolateral (al.w.) and posterolateral setal (pl.w.) warts oval (
Fig. 1C
). Maxillary palps five-segmented (
Fig. 1A, B
), labial palps three-segmented (
Fig. 1A
).
Thorax:
Prothorax with one pair of transversely elongate pronotal setal warts (
Fig. 1C
). Legs slender; tibial spur (sp.) formula 0, 1, 1–2, 2, 2 (e.g.
Fig. 1D–H
); hind tibia with 15–35 blackish spines (s.) (
Fig. 1G
). Forewing (Fig. 2A) narrow, slightly widening towards rounded apex, forks 1 and 5 present; discoidal cell (dc) about as long as thyroidal cell (tc). Hind wing (Fig. 2D) discoidal cell absent; crossvein r
1
–r
2
absent; crossvein r–m straight or convex; forks 1, 3, and 5, 1 and 5, or only 5, present; anterior edge with rasp-like, basal hairs (Fig. 2B) and 15–25 hamuli present past midway (Fig. 2C). Nygma (n): present in both wings in apical cell posterior to
R
4
(Fig. 2A, D).
Male genitalia (Fig. 2E–I):
Segment IX (t.IX and s.IX) annular, laterally setose, narrowest at base of superior appendages (s.app.) or ventrally; in lateral view tergite IX (t.IX) narrower to wider than sternite IX (s.IX) in lateral view. Superior appendages setose, flattened dorsoventrally, and as long as or longer than tergum X (t.X). Tergum X with median process bifurcated or entire along its length, membranous, and bearing small spines; pair of pre-apical, lateral, sclerotized processes (pl.p.) either absent or present. Inferior appendages bipartite, setose; basal part (if.b.) produced into posteromesal lip, with small, strong spines along dorsal margin; apical part of inferior appendage (if.ap.) long, digitate, with 25–60 mesal spines, mesal process with between four and nine mesal spines. Harpago (h.) absent or small. Phallic apparatus simple, phallicata basally tubular, fused with phallobase, curving to various degrees ventrad along its length; pair of sclerotized, ventrally produced lateral processes present apically; phallotremal sclerite (ph.sc.) U-shaped.
Inclusive species:
Gracilipsodes aoupiniensis
sp. nov.
;
Gracilipsodes aureus
sp. nov.
;
Gracilipsodes aurorus
sp. nov.
;
Gracilipsodes grandis
sp. nov.
;
Gracilipsodes koghiensis
sp. nov.
;
Gracilipsodes lanceolatus
sp. nov.
;
Gracilipsodes robustus
sp. nov.
;
Gracilipsodes psocopterus
Sykora (1967)
;
Gracilipsodes similis
Ward (2001)
.