An update of morphological and distributional data of the genus Patrera Simon (Araneae: Anyphaenidae: Anyphaeninae) with the description of twenty-five new species from Colombia Author Martínez, Leonel 0000-0002-4166-0561 leonelmarbio@gmail.com Author Brescovit, Antonio D. 0000-0002-1511-5324 antonio.brescovit@butantan.gov.br Author Villarreal, Eduardo 0000-0003-1303-2573 leonelmarbio@gmail.com Author Oliveira, Luiz Fernando M. 0000-0002-4054-4681 text Zootaxa 2021 2021-01-19 4914 1 1 64 journal article 8672 10.5281/zenodo.4468383 5c5c8aad-73f1-4a26-bdca-d1d02ba3fec1 4468383 Patrera carvalhoi new species Figures 15A–G ; 32 Type material. Male holotype from Quebrada Palo Blanco ( 4°43′40.08″N ; 75°34′48.36″W ), 1900m , Santuario de Fauna & Flora Otún Quimbaya , Vereda La Suiza, Corregimiento La Florida , Risaralda , Colombia , II-IV.2005, A. Sabogal leg. (ICN-Ar-10615). Paratypes : one female with same data (ICN-Ar-10631) . Etymology. The specific epithet is a patronym in honor of Leonardo Carvalho, for his contributions to the taxonomy and biogeography of Neotropical pholcids. Diagnosis. Males of Patrera carvalhoi n. sp. resemble those P. sampedroi n. sp. , by having the similar cymbium shape, S-shaped, long embolus, apically bilobed retrolateral tibial apophysis, and prominent tutacular groove on the cymbium ( Fig. 19 B–D), but can be diagnosed by the retrolateral tibial apophysis very short, entire and slightly bilobed in the apex (strongly lobulated in the apex, longer and wider in P. sampedroi ) ( Fig. 15 C–E). Females can be recognized from those of the remaining species of the genus by the copulatory ducts wide and coiled, and by the bean-shaped spermathecae ( Fig. 15 F–G). Description. Male ( Holotype , ICN-Ar-10615). Carapace brownish, darker on the cephalic region ( Fig. 15A ). Chelicerae dark brown. Labium and endites brown. Sternum yellow, darkest anteriorly. Legs brownish, metatarsus and tarsus darker. Abdomen yellow ( Fig. 15A ). Spinnerets yellow. Total length 8.76, carapace length 3.51, width 2.83, high 0.93. Clypeus height 0.2. Eye diameters and interdistances: AME 0.10, ALE 0.15, PME 0.21, PLE 0.21; AME–AME 0.29, AME–ALE 0.4, PME–PME 0.54, PME–PLE 0.49, ALE–PLE 0.46. Chelicerae 2.1 long, four promarginal teeth; four retromarginal teeth. Leg measurements: leg I—femur 5.91/ patella 1.94/ tibia 7.08/ metatarsus 5.47/ tarsus 2.57/ total 22.97; II—5.62/ 1.83/ 6.68/ 5.42/ 2.47/ 22.02; III—3.84/ 1.34/ 3.55/ 3.93/ 1.65/ 14.31; IV—5.14/ 1.49/ 4.58/ 5.9/ 1.67/ 18.78. Leg spination: I— tibia d1-0-1, metatarsus p1-1-0, r1-1-0; II— tibia d1-0-1, metatarsus p1-1-0, r1-1-0; III— tibia d1-0-1, v2-2-0, p0-1-1, r0-1-1, metatarsus p1-2-2, r1-1-2; IV— tibia d1-0-1, p0-1-1, r0-1-1, metatarsus p1-2-2, r1-1-2. Abdomen: length 5.21, epigastric furrow 1.21 from tracheal spiracle, spiracle 3.27 from base of spinnerets. Palp: retrolateral tibial apophysis short, small and apically bilobed; cymbium as long as half tibia length; subtegulum longer than wide, thin, with the distal projection inconspicuous; tegulum longer than wide; median apophysis long, laminar and medially situated; embolus long, apically filiform with narrow base and basally inserted on the tegulum ( Fig. 15 C–E). FIGURE 15A–G. Patrera carvalhoi n. sp. Male (ICN-Ar-10615): A habitus, dorsal view; C left palp, retroventral view; D palp, ventral view; E palp, retrolateral view. Female (ICN-Ar-10631): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Scale bars: A–B: 2mm; C: 0.8; D–E: 0.5mm; F–G: 0.2mm. Female ( Paratype , ICN–Ar 10631). Carapace pale brownish, with patches of dark brown setae along midline, ocular area darkest ( Fig. 15B ). Chelicerae brownish. Labium and endites brownish. Sternum yellow, margin darker, anteriorly lightest. Legs yellow basally, becoming brownish with some brown spots from distal side of femora to the patellae, from tibiae to tarsi yellow with some brown strips. Abdomen dorsally yellow with some patches of dark brown setae ( Fig. 15B ); ventrally yellow with some gray spots. Spinnerets yellow ( Fig. 7B ). Total length 7.95, carapace length 2.71, width 2.36, high 1.08. Clypeus height 0.11. Eye diameters and interdistances:AME 0.09, ALE 0.14, PME 0.17, PLE 0.19; AME–AME 0.26, AME–ALE 0.29, PME–PME 0.42, PME–PLE 0.42, ALE–PLE 0.39. Chelicerae 1.46 long, five promarginal teeth; four retromarginal teeth. Leg measurements: leg I—femur 3.7/ patella 1.34/ tibia 4.4/ metatarsus 3.04/ tarsus 1.36/ total 13.84; II—3.64/ 1.35/ 3.44/ 2.99/ 1.56/ 12.98; III—2.68/ 1.09/ 2.31/ 2.51/ 0.98/ 9.57; IV—3.64/ 1.08/ 3.1/ 3.52/ 1.08/ 12.42. Leg spination: I—metatarsus v2-0-0, p1-1-0, r1-1-0; II— tibia r1-0-1; metatarsus v2-0-0, p1-1-0, r1-1-0; III— tibia d1-0-1, v2-2-1, p0-1-1, r0-1-1, metatarsus p1-2-2, r1- 1-2; IV— tibia d1-0-1, p0-1-1, r0-1-1, metatarsus p1-2-2, r1-1-2. Abdomen: length 5.13, epigastric furrow 1.2 from tracheal spiracle, spiracle 2.57 from base of spinnerets. Epigynum: hood triangular; lateral borders weakly sclerotized, thin, semicircular; atrium not delimited by the lateral borders; internally with short, sinuousus copulatory ducts; seminal receptacles small, inserted on the anterior third of the copulatory ducts, spermathecae small, slender, with an anterior extension and posteriorly positioned; fertilization ducts shorter than spermathecae ( Fig. 15 F–G). Distribution. Only known from Risaralda department ( Fig. 32 )