An update of morphological and distributional data of the genus Patrera Simon (Araneae: Anyphaenidae: Anyphaeninae) with the description of twenty-five new species from Colombia Author Martínez, Leonel 0000-0002-4166-0561 leonelmarbio@gmail.com Author Brescovit, Antonio D. 0000-0002-1511-5324 antonio.brescovit@butantan.gov.br Author Villarreal, Eduardo 0000-0003-1303-2573 leonelmarbio@gmail.com Author Oliveira, Luiz Fernando M. 0000-0002-4054-4681 text Zootaxa 2021 2021-01-19 4914 1 1 64 journal article 8672 10.5281/zenodo.4468383 5c5c8aad-73f1-4a26-bdca-d1d02ba3fec1 4468383 Patrera wiwa new species Figures 21A–G ; 32 Type material. Male holotype from San Pedro Carmelo ( 10°52′N ; 73°43′W ), 1311m , Sierra Nueva Granada , Sierra Nevada de Santa Marta , Magdalena , Colombia , 24.IV.1975 , J. A. Kochalka leg. ( IBSP 213419 ). Paratypes : one male and one female from Sierra Nueva Granada, ( 10°52′N ; 73°43′W ), 1311m , Sierra Nevada de Santa Marta , Magdalena, Colombia , 24.IV.1975 , J. A. Kochalka leg. ( IBNP ) . Other material examined. COLOMBIA , Magdalena : Sierra Nevada de Santa Marta , Sierra Nueva Granada , San Pedro Carmelo ( 10°52′N ; 73°43′W ), 4100m , ♀ 1, 7.II.1974 , J. A. Kochalka leg. ( IBSP 213420 ) . Etymology. The specific name is a noun in apposition honoring to the Wiwa, one of the four Indian tribes that live on the slopes of the Sierra Nevada de Santa Marta. Diagnosis. Males of Patrera wiwa n. sp. resemble those of P. florezi n. sp. and P . yukpa n. sp. by the similarly shaped cymbium S-shaped, long embolus, and bifid retrolateral tibial apophysis ( Figs 16 C-E, 20B–D), but can be diagnosed by their short retrolateral tibial apophysis with the ventral branch shorter than the dorsal one (longer in P. florezi and almost with same length in P. yukpa ), median apophysis short, slender projected retrolaterally (larger and hook-shaped in P. florezi and P. yukpa ) ( Fig. 21 C–E). Females can be distinguished from those of the remaining species of the genus by the long, and thin hood, large copulatory ducts, and rounded and conspicuous seminal receptacles close to the spermathecae ( Fig. 21 F–G). FIGURE 21A–G. Patrera wiwa n. sp. Male (IBSP 213419): A habitus, dorsal view; C left palp, retroventral view; D palp, ventral view; E palp, retrolateral view. Female (IBSP 213419): B habitus, dorsal view; F epigynum, ventral view; G epigynum, dorsal view. Scale bars: A–B: 3.5mm; C, E: 0.73mm; D: 0.83mm; F–G: 0.25mm. FIGURE 22A–D. Patrera armata ( Chickering, 1940 ) . Male (ICN-Ar-10608): A habitus, dorsal view; B Chelicerae ventral view; C left palp, retroventral view (Arrow: indicates projection basal of the tibia); D palp, ventral view; E palp, retrolateral view. Female (ICN-Ar-5481): B habitus, dorsal view; G epigynum, ventral view; F epigynum, dorsal view. Abbreviations: BE, base of the embolus; Cvp, cheliceral ventral projection; dps, dorsal projection of the subtegulum; E, embolus; FD, fertilization ducts; H, hood; MA, median apophysis; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; ST, subtegulum; T, tegulum. Scale bars: A–B: 2mm. C: 1mm; D–F: 0.5mm; G–H: 0.25mm. FIGURE 23A–E. Patrera sutu n. sp. Male (ICN-Ar-10601): A habitus, dorsal view; B chelicerae, ventral view (Arrow: indicates projection of the chelicerae); C palp, ventral view (Arrow: indicating the left palp ventral view); D palp, retroventral view; E palp, ventral view. Abbreviations: E, embolus; Cvp, cheliceral ventral projection; MA, median apophysis; RTA, retrolateral tibial apophysis; SD, spermatic ducts; ST, subtegulum; T, tegulum; VTP, ventral tegular process. Scale bars: A: 2mm; B: 1mm; C: 0.2mm; D–E: 0.5mm. Description. Male ( Holotype , IBSP 213419). Cephalothorax orange, with dorsal darker paramedian bands ( Fig. 21A ). Legs yellow, ventrally orange. Abdomen dorsally cream with dispersed spots, more agglutinated medially; ventrally white ( Fig. 21A ). Total length 5.8, carapace length 2.7, width 2.5. Clypeus height 0.10. Eye diameters and interdistances: AME 0.06, ALE 0.14, PME 0.12, PLE 0.14; AME–AME 0.04, AME–ALE 0.02, PME–PME 0.14, PME–PLE 0.06, ALE–PLE 0.04. Chelicerae 1.1 long, four promarginal teeth; four retromarginal denticles. Leg measurements: leg I — femur 3.2/ patella 1.2/ tibia 3.7/ metatarsus 3.1/ tarsus 1.4/ total 12.6; II—3.1/ 1.2/ 3.4/ 3.0/ 1.3/ 12.0; III—2.22/ 0.9/ 1.8/ 2.3/ 0.8/ 8.02; IV—3.0/ 1.0/ 2.8/ 3.43/ 1.0/ 11.23. Leg spination: I—tibia v2-2-2, p1-1-1, r1-1-1, metatarsus v2-1-0, p1-1-0, r1-1-0; II—tibia v2-2-2, p1-1-1, r1-1-1, metatarsus v2-1-0, p1-1-0, r1- 1-0; III—tibia v2-2-2, p0-1-1, r1-0-1-1, metatarsus v2-2-2, p1-1-1, d-1-1, r1-1-1; IV—tibia v2-2-2, p1-1-1-1, r1-1- 1-1, metatarsus v2-2-2, p1-1-1-1, r1-1-1-1-1. Abdomen: length 3.1, epigastric furrow 0.95 from tracheal spiracle, spiracle 1.3 from base of spinnerets. Palp: retrolateral tibial apophysis bifid, long, with the ventral branch shorter than the dorsal one; cymbium shorter than tibia, with protuberant tutacular groove; subtegulum longer than wide, with conspicuous distal projection; tegulum longer than wide, with a small, sclerotized ventral tegular process under the median apophysis; median apophysis short, curved and medially situated; embolus long, apically filiform, with narrow base and basally inserted on the tegulum ( Fig. 21 C–E). Female ( Paratype , IBSP 213419). Coloration as in the male, except dorsum of the abdomen with dark brown dorsal spots on medial side ( Fig. 21B ). Total length 8.0, carapace length 3.5, width 2.7. Clypeus height 0.14. Eye diameters and interdistances: AME 0.06, ALE 0.14, PME 0.14, PLE 0.10; AME–AME 0.06, AME–ALE 0.06, PME–PME 0.10, PME–PLE 0.18, ALE–PLE 0.06. Chelicerae 1.3 long; four promarginal teeth; six retromarginal denticles. Leg measurements: leg I—femur 3.5/ patella 1.3/ tibia 3.62/ metatarsus 2.9/ tarsus 1.4/ total 12.72; II— 3.5/ 1.3/ 3.4/ 2.8/ 1.3/ 12.3; III—2.4/ 1.1/ 1.7/ 2.3/ 0.8/ 8.3; IV—3.0/ 1.0/ 3.0/ 3.4/ 10.4. Leg spination: I—tibia v2-2-2, p1v-1-1-0, r1v-1-10, metatarsus v2-0-0, p1-1-0, r1-1-0; II—tibia v2-2-2, p1-1-1-0, r1-1-1-0, metatarsus v2- 2-2, p1-1-0, r1-1-0; III—tibia v2-2-2, p1-1-0, r1-1-0, metatarsus v2-2-2, p1-1-1, r1-1-1; IV—tibia v2-2-2, p1-1-1-0, r1-1-1-0, metatarsus v2-2-2, p1-1-1-1-1, r1-1-1. Abdomen: length 4.5, epigastric furrow 1.0 from tracheal spiracle, spiracle 2.1 from base of spinnerets. Epigynum: hood long and thin; lateral borders sclerotized, thin, oblique; atrium long, triangular; internally with long, n-shaped copulatory ducts; seminal receptacles small, rounded, medially situated on the copulatory ducts; spermathecae small, rounded, posteriorly positioned; fertilization ducts shorter than spermathecae length ( Fig. 21 F–G). Distribution. Only known from Magdalena department ( Fig. 32 ).