Pamphagidae (Orthoptera: Acridoidea) from the Palaearctic Region: taxonomy, classification, keys to genera and a review of the tribe Nocarodeini I. Bolívar Author Ünal, Mustafa text Zootaxa 2016 4206 1 1 223 journal article 37403 10.11646/zootaxa.4206.1.1 5c5bff7d-ea8c-4893-b03e-a8b5c5904514 1175-5326 208260 E1566C02-9987-4116-83AA-91D3D1DCF2FF Paranocarodes tolunayi tolunayi Ramme, 1949 ( Figs. 45 , 417–418 , 840 ) Paranocarodes Tolunayi Ram (n.sp.): in Karabağ 1949 : 66 , 102. Paranocarodes tolunayi n. sp. : Ramme 1951 : 278 , 282, 427. Paranocarodes tolunayi Karabağ, 1949 : Karabağ 1958 : 117 ; Weidner 1969 : 158 ; Presa & Garcia 1983 : 24 . Paranocarodes atympanicus n. sp. : Ramme 1951 : 278 , 285, 427, partim (female nymph holotype ). syn. nov. Paranocarodes atympanicus Ramme, 1951 : Karabağ 1958 : 117 ; Weidner 1969 : 158 ; Presa & Garcia 1983 : 24 ; Otte 1994 : 184 . Paranocarodes fieberi tolunayi Karabağ, 1949 : Demirsoy 1973 : 413 , partim; Demirsoy 1977 : 65 , partim; Otte 1994 : 184 ; Ünal 1999 : 247 . Paranocarodes tolunayi Ramme, 1951 : Otte 1994 : 185 . Type locality. Turkey , Ankara , Kızılcahamam , Gökdere. Holotype : male ( MfN ). Material examined. TURKEY : Anatolien , [ Ankara ], Kisilcahamam [ Kızılcahamam ], Gökdere , 25.5.1940 , 1♂ ( Holotype ), 1♂ , 1♀ nymph ( Paratypes ) (leg. T. Karabağ ) ( MfN ) ; Angora [ Ankara ], 1♀ nymph ( Holotype of Paranocarodes atympanicus ) (leg. Wiedemann ) ( MfN ) ; Ankara , Kızılcahamam , 25.5.1940 , 1♂ , 1♀ nymph, 25.4.1940 , 1♀ nymph [ Topotypes ] (leg. T. Karabağ ) ( NHMUK ) ; Ankara , Kızılcahamam , Gökdere , 25.5.1940 , 1♂ ( Paratype ) (leg. T. Karabağ , det. Ramme as Paranocarodes n.sp. ); ibidem 1♂ , 3♀ nymphs [ Topotypes ] (leg. T. Karabağ ) ( AÜZM ) ; Bolu , Kındıra Köyü , Güvenlik Yaylası , 1570 m , 1.7.2000 , 1♂ , 3♀ ; Ankara , Kızılcahamam , Güvem Köyü , 1100 m , 28.5.1997 , 1♂ (leg. M. Akçay ) ( Ünal , 2009 ) ; Bolu , Gerede , Naldöken Tepesi , 1900, 4.6.2008 , 1♂ , 1♀ (leg. M. Ünal ) ; Ankara , Kızılcahamam , Soğuksu Milli Parkı , 9.6.1993 , 1♂ (leg. M. Ünal ) (AİBÜEM). Distribution. Known from N. Turkey : along the Köroğlu Mountains in Ankara and Bolu provinces ( Fig. 840 ). Remarks. The author and the date of this species were given as “ Karabağ, 1949 ” by Karabağ (1958) , Weidner (1969) , Demirsoy (1973 , 1977 ), Presa & Garcia (1983) , Otte (1994) , Ünal (1999) . But, Karabağ (1949: 66) gave this species for the first time as “ Paranocarodes tolunayi Ram (n.sp.)”. Karabağ also added a short note just below this species: “Identification and description of this species that found by us from 1600 m . height of the Kızılcahamam forests, were made by Ramme”. Therefore it is clear that the author of this species must be Ramme (in Karabağ, 1949 ). The holotype of P. atympanicus Ramme is a female nymph though Ramme (1951: 285) did not mention its nymphal state. There are also 1 female nymph of P. tolunayi in the MfN, 1 female nymph in the NHMUK and 3 female nymphs in the AÜZM. They all were compared in detail and concluded that they are certainly identical and probably are members of the same population. Besides, the only known species of Paranocarodes in Ankara province is P. tolunayi . Considering the similarities of the holotype (female nymph) of P. atympanicus it has to be considered synonym of P. tolunayi tolunayi . On the other hand the paratype (adult female) of P. atympanicus from Denizli Province is P. karabagi (see P. karabagi above).