Australodindymus nigroruber gen. et sp. nov. from Western Australia (Hemiptera: Heteroptera: Pyrrhocoridae) Author Stehlík, Jaroslav L. Author Jindra, Zdenĕk text Zootaxa 2012 3316 57 62 journal article 10.5281/zenodo.281111 19e23780-6b44-4704-83ca-02cfee9acaef 1175-5326 281111 Australodindymus gen. nov. Type species. Australodindymus nigroruber sp. nov. , here designated. Description. Body dorsally matt, elongate, with costal margin of hemelytra slightly rounded (Fig. 1). Head prognathous, slightly wider than long. Eyes slightly convex, temples regularly rounded, eye sockets not developed, eyes adjacent the anterior margin of pronotum. Vertex wide, reaching base of eye temple. Frons slightly gibbous, elevated above eye level in lateral view. Clypeus in lateral view distinctly elevated above paraclypei; ventral side of head regularly rounded. Bucculae rounded. Labium of male reaching anterior margin of ventrite IV, in female somewhat shorter. Anterior margin of pronotal collar concave. Callar lobe slightly gibbous, as well as the pronotal lobe towards the base of pronotum. Base of scutellum convex, disc of scutellum slightly convex, apex elongated and pointed. Membrane slightly surpassing apex of abdomen; veins arising from closed cells formed by cubitus, media and radius sector, little branching and nearly reaching posterior margin of membrane. Ventral side of profemora rounded, apically concave and bearing three to four teeth. Apical halves of tibiae ventrally with obliquely erect spines; shorter stout setae also present on ventral side of tarsomere 1. Protarsus more than twice as long as meso- and metatarsus. Intersegmental sutures between abdominal ventrites slightly S-shaped laterally. In males ventrite VII medially wide and ventrite VI medially narrowed; in females ventrites VI and VII simple. Pygophore ( Figs. 3–7 ). Ventral two thirds of ventral wall rounded in lateral view, the dorsal third strongly produced posteriorly. Upper margin of ventral portion of the ventral wall laterally with short furrow. Ventral rim with rectangular incision. Ventral rim infolding straight, not rounded, sloping upright into genital chamber. Lateral rim sharp, regularly rounded, smoothly merging with the dorsal rim. Lateral rim infolding regularly rounded, strongly sloping into genital chamber. Pygophore dorsally with pale pilosity. Paramere (Fig. 8) slender in basal half, but strongly enlarged in distal half, apex rounded, and ventral surface of paramere strongly concave. Both parameres in situ distant, their apices facing each other. Female genitalia (Fig. 9). Both sides of valvifer I considerably separated from each other, small, upper edge arcuate. Anal tube rather short, transversely positioned. Laterotergites IX plate-like, situated just beneath the anal tube; basally very close, diverging from the base up, their distal margins arcuately embracing large, black, stackshaped projections of valvifer II. Valvifer II entirely exposed; another two black, stack-shaped projections present on dorsal part of valvifer II, closer to one another than those in ventral part. Differential diagnosis. The species-rich genus Dindymus is currently divided into five subgenera ( Dindymus s. str. , Anthridindymus Stehlík, 2006 , Cornidindymus Stehlík, 2005 , Limadindymus Stehlík, 2005 , Pseudodindymus Stehlík, 2009 ), differing especially by various modifications of parameres. The basic character of Dindymus is parameres crossing each other. The parameres could be attenuated distally with pointed apex, or very slender, rodshaped, distally slightly widened with rounded apex, or slightly curved anteapically. In such cases the paramere is simple, not differentiated into body and processus hamatus. By comparison, in Pseudodindymus the parameres are strongly reduced, their body is short, with an acicular processus hamatus, oriented towards the centre of the pygophore but not reaching it and therefore not crossing each other (see Stehlík 2009 : Figs. 7 –11). As an exception, in D . ( Pseudodindymus ) limbaticollis Breddin, 1901 only the body of paramere is developed ( Stehlík 2009 ). Australodindymus gen. nov. differs from all Dindymus species by the paramere being slender in basal half, but strongly enlarged in distal half, having rounded apex, and ventral surface of the paramere being strongly concave (Fig. 8). Both parameres in situ are distant from each other, their apices facing each other. In females of all subgenera of Dindymus laterotergite IX is large, medially longitudinally concave but never very thin, plate-like, with both sides approaching each other, as in Australodindymus gen. nov. Etymology. The generic name is composed from the prefix Australo -, meaning southern or Australian, and the generic name Dindymus , and refers to both the similarity of habitus of the new genus with Dindymus and its distribution in Australia . Gender is masculine.