Phylleremus n. gen., from leaves of deciduous trees in eastern Australia (Oribatida: Licneremaeoidea)
Author
Behan-Pelletier, Valerie M.
Author
Walter, David E.
text
Zootaxa
2007
1386
1
17
journal article
10.5281/zenodo.175132
8dacef77-cc64-40d3-92ad-be6e928111e2
11755326
175132
Phylleremus
n. gen.
Typespecies:
Phylleremus leei
n. sp.
Diagnosis.
Species in this genus have the following unique combination of character states: adult with tuberculate, nonbirefringent cerotegument; taenidium laterally on prodorsum; notogaster with one or four pairs of minute saccules, saccule Sa positioned on anterior margin of notogaster, far anterior of seta
c
; nine pairs of notogastral setae; posterior notogastral tectum present, with posteromedial incision covered by pair of overlapping lobes; palpal eupathidium
acm
attached to solenidion along length; two pairs of adanal setae; immatures apheredermous, plicate; seta
p3
absent from both immatures and adults; seta
d
absent from genua and tibiae in immatures and adults; femora, tibiae and tarsi with ridges running dorsoventrally in nymphs and adults; tarsi I to IV of immatures and adults with large, subunguinal pulvillus.
FIGURES 1–3.
Phylleremus leei
n. sp.
, adult female: 1, dorsal aspect; 2, ventral aspect (legs removed); 3, prodorsum, lateral aspect (gnathosoma, notogaster and legs removed; taenidium indicated by arrow).
Description
.
Adult:
Body and legs with weakly developed, tuberculate, nonbirefringent cerotegument. Integument with microtubercles. Dorsophragmata and pleurophragmata present, dorsophragmata positioned medially (
Fig. 1
). Lamellae and genal notch absent. Bothridial seta capitate (
Figs 1
,
6
). Bothridium small, with internal ringlike ridges (
Fig. 3
). Porose area Ad posterolateral to seta
in
(
Fig. 3
). Pedotecta I and II covering base of acetabula I and II, respectively (
Fig. 3
). Taenidium extending from seta
ex
to edge of rostrum (
Figs 3
,
6
). Pedotectum I extending dorsally to base of seta
ex
(
Fig. 3
). Coxisternal setation: 3122 (
Fig. 2
). Apodemes 2 and sejugal apodeme almost transverse. Custodium absent, discidium developed as tubercle between acetabula III and IV (
Fig. 2
). Tracheal system normal. Genital setae 5 or 6 pairs. Aggenital setae 1 pair. Anal setae 2 pairs. Adanal setae 2 pairs. Lyrifissure
ian
absent. Preanal organ as caecum. Postanal porose area absent. Humerosejugal porose areas Am and Ah, absent. Sublamellar porose area Al absent (
Fig. 3
). Notogaster fused with prodorsum medially, slightly flattened anteriorly, ahumerate (
Fig. 1
). Octotaxic system present as one or four pairs of saccules, Sa positioned near anterior edge of notogaster, anterior to seta
c,
and dorsal to dorsosejugal porose area Ad (
Figs 1
,
19
) Nine pairs of notogastral setae present,
c2
, and
l
and
h
series positioned laterally; setae
p1
and
p2
positioned marginally (
Fig. 2
);
c1, c3, da, dm, dp,
and
p3
absent. Notogastral tectum present, indented and overlapping posteromedially (
Fig. 7
). Lyrifissures
ia
positioned slightly posterolaterally of setae
c2
,
im
positioned between setae
lm
and
lp
;
ih
positioned lateral of setae
h1
and
p1
,
ip
and
ips
positioned laterally on notogaster anterior to seta
p2
. Pteromorphs absent. Subcapitulum diarthric, axillary saccule absent from base of palp. Rutellum pantelobasic. Chelicera chelatedentate with 2 slender, barbed setae. Trägårdh’s organ present. Palp with normal 5 segments and tarsal lyrifissure; setal formula: 0213 9(1). Eupathidium
acm
fused with solenidion along length (
Fig. 17
). Legs monodactyl, tarsi with subunguinal pulvillus (
Figs 15, 16
). Femora I to IV and trochanters III and IV with porose areas (
Figs 8–11
). Famulus
(e)
rodlike, rounded distally. Seta
d
absent from tibiae I to IV and genua I to III. Solenidion ω2 present on tarsus II. Solenidia on tibiae and genua short, other than φ
1
on tibia I. Tibia I without tubercle projecting over base of tarsus I.
Immatures:
Apheredermous, plicate, without hysterosomal macrosclerites or excentrosclerites. Line of dehiscence extends anterior to seta
c2
(
Fig. 12
). Cerotegument tuberculate (
Fig. 18
). Integument smooth. Prodorsal porose regions present (
Fig. 12
). Gastronotal setation bideficient; larva with 12, nymphs with 14 pairs of setae (
f1
,
p3
absent). Gastronotal setae of
c
,
d
, and
l
series monomorphic in nymphs (
Figs 12
,
20
). Opisthonotal gland present in all instars. Coxisternal porose regions present in all nymphs (
Fig. 13
). Porose regions present lateral of genital region, in adanal region, and surrounding opening of opisthonotal gland in all nymphs (
Fig. 13
; porose regions lateral of genital region not illustrated). Apodematoacetabular tracheal system or porose homologues absent. Paraprocts atrichous in larva, protonymph and deutonymph. Genital and aggenital setal formula (larva to adult): 01356 or 01?45, and 00111, respectively. Cupule development normal. Bothridium, bothridial seta and seta
in
fully formed in all immatures. Bothridium cupshaped. Seta
d
absent from tibiae I to IV and genua I to III. Setation of protonymphal leg IV normal: 00007.
Etymology.
The generic prefix “Phyll” is from the Greek “phyllon” meaning leaf, and refers to the leaf habitat of members of this genus; “eremus” is a common suffix for oribatid genera and is from the Greek “eremos’ meaning solitary.