Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group Author Golovatch, Sergei I. Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru sgol@orc.ru Author Geoffroy, Jean-Jacques Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr geoffroy@mnhn.fr Author Mauriès, Jean-Paul Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr collmill@mnhn.fr Author Spiegel, Didier Van Den Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be text Zoosystema 2007 29 1 7 49 journal article 10.5281/zenodo.5392079 1638-9387 5392079 Glyphiulus paragranulatus n. sp. ( Figs 10-12 ) TYPE MATERIAL. — China . Yunnan Prov. , Jianshui County , Yan Dong Cave , 12.I.1989 , leg. P. Beron , holotype ♂ ( NMNHS ) ; paratypes 2 ♀♀ , 1 fragm. juv. ( NMNHS ) ; 1 ♂ , 1 ♀ ( MNHN GA 036 ) ; 1 ♂ ( ZMUM ) ; 1 ♂ , 1 ♀ ( SEM ) . ETYMOLOGY. — To emphasize the obvious similarity to G. granulatus . DIAGNOSIS. — Differs from the most similar congeners by the antennae being more slender, the body segments somewhat compressed laterally, the ocellaria only slightly reduced, the accessory spine at the claw base especially prominent, the anterior gonopod coxosternum with small but evident distomedial outgrowths. DESCRIPTION Length of adults of both sexes 18-25 mm , width 1.0- 1.2 mm . General coloration pallid, pattern much as in G. granulatus , but ocellaria light brown while lateral stripes level to ozoporiferous tubercles narrow. Adults with 39-45p+2-1a+T. Holotype about 25 mm long, 1.2 mm wide, with 44p+2a+T. Other characters as in G. granulatus , including gnathochilarium (divided into pro- and eumentum, n = 1) ( Fig. 12B ), the carinotaxy formulae ( Figs 10 A-D; 11B), telson ( Figs 10E, F ; 11A ), male legs 1-3 ( Fig. 12 C-E), posterior gonopod structure ( Figs 11E ; 12G, H ), etc., but body size much larger (see above); segments compressed laterally ( Fig. 11B ); antennae slightly elongate ( Figs 10A, B ; 12A ); claw with a particularly prominent spine at base ( Fig. 11D ); only some 9-11 ocelli in adults; legs somewhat longer (about as long as body height) ( Figs 10D ; 11C ); anterior gonopod coxosternum with small but curved distomedian outgrowths and acute distolateral corners ( Figs 11F ; 12F ). REMARKS Even though there are some indications of troglomorphism in this species (pallid general coloration, somewhat elongated antennae and legs), its troglophily seems more likely than troglobiosis. The morphological evidence being inconclusive, without further observations this problem can never be solved. On the other hand, Bollmania beroni Stoev & Enghoff, 2005 ( Diplopoda , Callipodida , Caspiopetalidae Lohmander,1931 ), a presumed troglobitic millipede, occurs in the same cave ( Stoev & Enghoff 2005 ).