Laboulbeniomycetes (Fungi, Ascomycota) of Denmark
Author
Santamaria, Sergi
Unitat de Botànica, Departament de Biologia Animal, de Biologia Vegetal i d’Ecologia, Facultat de Biociències, Universitat Autònoma de Barcelona, 08193 Cerdanyola del Vallès (Barcelona), Spain.
sergi.santamaria@uab.cat
Author
Pedersen, Jan
Natural History Museum of Denmark (Zoological Museum), University of Copenhagen, Universitetsparken 15, 2100 København Ø, Denmark.
japedersen@snm.ku.dk
text
European Journal of Taxonomy
2021
2021-11-26
781
1
425
http://dx.doi.org/10.5852/ejt.2021.781.1583
journal article
10.5852/ejt.2021.781.1583
2118-9773
5828924
Laboulbenia pedicellata
Thaxt.
MB#219334
Fig. 40C–H
Proceedings of the American Academy of Arts and Sciences
27: 44 (
Thaxter 1892
)
. –
Type
: “
On
Bembidium
sp.
,
Maine
”;
FH
. [
USA
]
Laboulbenia gracilipes
Cépède & F.Picard. (
Cépède & Picard 1907: 780
)
[MB#182939]
Laboulbenia gibbulosa
Speg. (
Spegazzini 1915b: 55
)
[MB#183213]
Laboulbenia tapirina
R.K.Benj. & Shanor (
Benjamin & Shanor 1952: 126
)
[MB#299202]
Diagnostic features
Cell V variably oval, as long as cell IV, connected with septum III–IV. Outer appendage variably branched above the suprabasal cell; septa at the lower side in outer branch thick and dark, with blackened suffusion spreading through the dorsal margin (
Fig. 40G
, arrow). Inner appendage consisting of numerous branches bearing lateral solitary antheridia. Basal cell of inner appendage slightly shorter than basal cell of outer appendage. [Detailed descriptions:
Thaxter 1896
;
Huldén 1985
;
Majewski 1994b
;
Santamaria 1998
]
Distribution and hosts
Common on many species of the genus
Bembidion
in a broad sense, and on
Dyschirius
(Col.
Carabidae
). Occurrence on other hosts are likely wrong or require verification. Widespread, only unknown in Oceania (
Santamaria
et al.
1991
;
Majewski 2008
). Recent records are from
Denmark
(
Høier 2009
),
Switzerland
(
Hoess & Senn-Irlet 2009
),
Slovakia
(
Rossi
et al.
2010
),
Sweden
(
Huggert 2010
),
the Netherlands
(
Haelewaters
et al.
2012
),
Israel
,
Ecuador
(
Barragán
et al.
2013
),
Bulgaria
,
Cyprus
(
Rossi
et al.
2019a
),
Ukraine
(
Haelewaters
et al.
2019b
), and
Cambodia
(
Kong
et al.
2020
).
Fig. 40.
Laboulbenia
spp.
A
–B
.
L. parriaudii
Balazuc ex Santam. Mature
thalli showing the distal septum of the basal cell darkened and constricted (arrows), a diagnostic character. –
C–H
.
L. pedicellata
Thaxt. Mature
thalli, except the immature one represented in G showing the septum at the lower side in outer branch thick and dark, with blackened suffusion spreading through the dorsal margin (arrow).
C–D
. From
Bembidion aeneum
.
E
. From
Bembidion doris
.
F
. From
Bembidion minimum
.
G–H
. From
Bembidion mannerheimi
. Scale bars: 50 µm. Photographs from slides ZMUC C-F-124096 (A–B), ZMUC C-F-122506 (C–D), ZMUC C-F-122492 (E), ZMUC C-F-123118 (F), ZMUC C-F-123132 (G), ZMUC C-F-123391 (H).
Collections examined from
Denmark
On
Bembidion aeneum
Germar, 1824
(Col.
Carabidae
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Resle Skov
;
54°56.359′ N
,
11°43.437′ E
; PF79;
25 Sep. 2006
;
H. Liljehult
1461
;
JP
det.;
ZMUC C-F-124012
. –
Nordøstsjaelland
(NEZ)
•
Eriksholm Skov
;
55°41.771′ N
,
11°47.912′ E
; PG77;
4 Nov. 2017
;
JP
740
;
JP
det.;
ZMUC C-F-123234
•
Nord
for
Frederikssund
;
55°51.404′ N
,
12°3.534′ E
; UB17;
14 Dec. 2013
;
R. Aagaard Jensen
409
;
JP
det.;
ZMUC C-F-122893
•
Nord
for
Risø
;
55°42.184′ N
,
12°6.692′ E
; UB17;
14 Dec. 2013
;
R. Aagaard Jensen
628
;
JP
det.;
ZMUC C-F-123117
. –
Nordvestjylland
(NWJ)
•
Nord
for
Landting
ved
Vinderup
;
56°30.049′ N
,
8°45.384′ E
; MH86;
19 Jan. 2015
;
JP
573
;
JP
det.;
ZMUC C-F-123059
•
Syd
for
Stårup
;
56°34.792′ N
,
9°5.958′ E
; NH07;
6 Feb. 2015
;
K.B. Nielsen
218
;
JP
det.;
ZMUC C-F-122699
•
Vest
for
Vinderup
;
56°28.727′ N
,
8°44.810′ E
; MH85;
19 Jan. 2015
;
JP
779
;
JP
det.;
ZMUC C-F-123274
. –
Nordvestsjaelland
(NWZ)
•
Bognaes Skov
på
Tuse Naes
;
55°44.966′ N
,
11°45.817′ E
; PG78;
10 Dec. 2013
;
JP
873
;
JP
det.;
ZMUC C-F-123369
•
ibid.;
30 Dec. 2013
;
K.B. Nielsen
30
;
JP
det.;
ZMUC C-F-122506
. –
Sydjylland
(SJ)
•
Ballum Sluse
;
55°7.866′ N
,
8°41.125′ E
; MG70;
26 Oct. 2017
;
H. Liljehult
872
;
JP
det.;
ZMUC C-F-123368
.
On
Bembidion assimile
Gyllenhal, 1810
(Col.
Carabidae
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Lysemose
ved
Maribo
;
54°46.971′ N
,
11°28.959′ E
; PF57;
4 Jun. 2018
;
JP
1128
;
JP
det.;
ZMUC C-F-123651
. –
Nordøstsjaelland
(NEZ)
•
Nordhavnstippen
på
Københavns Nordhavn
;
55°43.091′ N
,
12°37.088′ E
; UB57;
29 Apr. 2018
;
JP
1092
;
JP
det.;
ZMUC C-F-123609
•
Ud
for
Kongelundsfortet
;
55°33.724′ N
,
12°33.731′ E
; UB46;
15 Apr. 2018
;
JP
1056
;
JP
det.;
ZMUC C-F-123573
. –
Sydsjaelland
(SZ)
•
Gammel Kalvehave
;
54°59.745′ N
,
12°8.232′ E
; UA19;
25 Apr. 2019
;
JP
1448
;
JP
det.;
ZMUC C-F-123996
.
On
Bembidion biguttatum
(Fabricius, 1779)
(Col.
Carabidae
)
DENMARK
–
Bornholm
(B)
•
Skelsmyre
i
Rønne Plantage
;
55°4.375′ N
,
14°45.096′ E
; VB80;
25 Apr. 2014
;
JP
508
;
JP
det.;
ZMUC C-F-122991
. –
Lolland, Falster, Møn
(LFM)
•
Lysemose
ved
Maribo
;
54°46.971′ N
,
11°28.959′ E
; PF57;
4 Jun. 2018
;
JP
1129
;
JP
det.;
ZMUC C-F-123652
•
Nyord
;
55°2.885′ N
,
12°12.380′ E
; UB20;
27 Apr. 2013
;
JP
837
;
JP
det.;
ZMUC C-F-123332
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1322
;
JP
det.;
ZMUC C-F-123858
. –
Nordøstsjaelland
(NEZ)
•
Amager Faelled
;
55°38.631′ N
,
12°34.729′ E
; UB47;
22 Apr. 2018
;
JP
1038
;
JP
det.;
ZMUC C-F-123554
. –
Sydsjaelland
(SZ)
•
Østerskov
ved
Langebaek
;
54°59.572′ N
,
12°6.715′ E
; UA19;
3 Jan. 2019
;
JP
1287
;
JP
det.;
ZMUC C-F-123819
.
On
Bembidion doris
(Panzer, 1796)
(Col.
Carabidae
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Ravnsholt
;
54°57.415′ N
,
11°15.306′ E
; PF49;
24 Aug. 2018
;
L. Kjaer-Thomsen
1176
;
JP
det.;
ZMUC C-F-123700
•
Ulvshale
;
55°3.218′ N
,
12°15.801′ E
; UB20;
10 Jun. 2017
;
JP
429
;
JP
det.;
ZMUC C-F-122914
. –
Nordøstsjaelland
(NEZ)
•
Fruebjerg
i
Strøgårdsvang
;
55°58.539′ N
,
12°16.659′ E
; UC30;
18 Oct. 2018
;
JP
1223
;
JP
det.;
ZMUC C-F-123750
•
Indelukket
ved
Frederiksborg Slot
;
55°56.191′ N
,
12°17.861′ E
; UC30;
12 Apr. 2016
;
JP
19
;
JP
det.;
ZMUC C-F-122492
.
On
Bembidion gilvipes
Sturm, 1825
(Col.
Carabidae
)
DENMARK
–
Nordøstsjaelland
(NEZ)
•
Stålmosen
,
Vindinge
;
55°37.379′ N
,
12°8.322′ E
; UB16;
12 Apr. 1979
;
H.J. Petersen
Dry0055
;
H.J. Petersen
;
ZMUC C-F-124121
. –
Sydsjaelland
(SZ)
•
Østerskov
ved
Langebaek
;
54°59.572′ N
,
12°6.715′ E
; UA19;
3 Jan. 2019
;
JP
1288
;
JP
det.;
ZMUC C-F-123820
•
Tryggevaelde Å ved
Varpelev
;
55°21.055′ N
,
12°15.877′ E
; UB23;
6 Jan. 2018
;
JP
1589
;
JP
det.;
ZMUC C-F-124350
.
On
Bembidion guttula
(Fabricius, 1792)
(Col.
Carabidae
)
DENMARK
–
Nordøstsjaelland
(NEZ)
•
Nord
for
Risø
;
55°42.184′ N
,
12°6.692′ E
; UB17;
14 Dec. 2013
;
R. Aagaard Jensen
627
;
JP
det.;
ZMUC C-F-123116
•
Syd
for
Veddelev
;
55°40.346′ N
,
12°4.915′ E
; UB17;
14 Dec. 2013
;
R. Aagaard Jensen
333
;
JP
det.;
ZMUC C-F-122811
. –
Nordvestjylland
(NWJ)
•
Nord
for
Landting
ved
Vinderup
;
56°30.049′ N
,
8°45.384′ E
; MH86;
19 Jan. 2015
;
JP
574
;
JP
det.;
ZMUC C-F-123060
•
Vest
for
Vinderup
;
56°28.727′ N
,
8°44.810′ E
; MH85;
19 Jan. 2015
;
JP
777
;
JP
det.;
ZMUC C-F-123272
. –
Nordvestsjaelland
(NWZ)
•
Flasken
nord for
Reersø
;
55°31.965′ N
,
11°9.831′ E
; PG35;
26 Jan. 2014
;
JP
611
;
JP
det.;
ZMUC C-F-123099
•
Halleby Å ved
Bjerge Sydstrand
;
55°33.194′ N
,
11°10.079′ E
; PG35;
24 Oct. 2018
;
JP
1211
;
JP
det.;
ZMUC C-F-123738
. –
Sydjylland
(SJ)
•
Brede Ådal
;
55°3.184′ N
,
8°54.138′ E
; MG90;
24 Oct. 2013
;
H. Liljehult
500
;
JP
det.;
ZMUC C-F-122984
. –
Sydsjaelland
(SZ)
•
Even Bro
;
55°8.694′ N
,
12°0.601′ E
; UB11;
17 Feb. 2019
;
JP
1370
;
JP
det.;
ZMUC C-F-123911
•
Tryggevaelde Å ved
Varpelev
;
55°21.055′ N
,
12°15.877′ E
; UB23;
6 Jan. 2018
;
JP
967
;
JP
det.;
ZMUC C-F-123474
.
On
Bembidion illigeri
Netolitzky, 1914
(Col.
Carabidae
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Rødbyhavn
;
54°39.072′ N
,
11°21.809′ E
; PF55;
4 Jun. 2018
;
JP
1120
;
JP
det.;
ZMUC C-F-123640
.
On
Bembidion lunulatum
(GeoffroY, 1785)
(Col.
CArABIdAe
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1323
;
JP
det.;
ZMUC C-F-123859
. –
Nordøstsjaelland
(NEZ)
•
Brønshøj
(Degnemosen);
55°42.043′ N
,
12°30.352′ E
; UB47;
26 May 2016
;
JP
290
;
JP
det.;
ZMUC C-F-122769
.
On
Bembidion mannerheimii
Sahlberg, 1827
(Col.
Carabidae
)
DENMARK
–
Fyn
(F)
•
Gyldenstens
inddaemmede
Strand
;
55°34.447′ N
,
10°7.211′ E
; NG75;
2 Apr. 2014
;
JP
710
;
JP
det.;
ZMUC C-F-123200
•
Kohaveskov
ved
Landkildegård
;
55°22.750′ N
,
10°27.979′ E
; NG93;
24 Aug. 2001
;
H. Liljehult
1398
;
JP
det.;
ZMUC C-F-123942
,
C-F-123944
. –
Lolland, Falster, Møn
(LFM)
•
Krenkerup Haveskov
;
54°46.408′ N
,
11°39.955′ E
; PF77;
23 Sep. 1993
;
S. Langemark
&
O.E. Meyer
893
;
JP
det.;
ZMUC C-F-123391
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1324
;
JP
det.;
ZMUC C-F-123863
. –
Nordøstjylland
(NEJ)
•
Syd
for
Krogsagergård
;
56°37.534′ N
,
9°45.551′ E
; NH47;
24 Nov. 2019
;
JP
1555
;
JP
det.;
ZMUC C-F-124316
. –
Nordøstsjaelland
(NEZ)
•
Eriksholm Skov
;
55°41.771′ N
,
11°47.912′ E
; PG77;
4 Nov. 2017
;
JP
741
;
JP
det.;
ZMUC C-F-123235
. –
Nordvestsjaelland
(NWZ)
•
Vesterlyng
;
55°44.195′ N
,
11°17.276′ E
; PG47;
9 Feb. 2014
;
JP
644
;
JP
det.;
ZMUC C-F-123132
. –
Sydsjaelland
(SZ)
•
Gammel Kalvehave
;
54°59.745′ N
,
12°8.232′ E
; UA19;
25 Apr. 2019
;
JP
1449
;
JP
det.;
ZMUC C-F-123999
•
Østerskov
ved
Langebaek
;
54°59.235′ N
,
12°6.280′ E
; UA19;
14 Jan. 2012
;
JP
546
;
JP
det.;
ZMUC C-F-123032
.
On
Bembidion minimum
(Fabricius, 1792)
(Col.
Carabidae
)
DENMARK
–
Nordøstsjaelland
(NEZ)
•
Nord
for
Risø
;
55°42.184′ N
,
12°6.692′ E
; UB17;
14 Dec. 2013
;
R. Aagaard Jensen
629
;
JP
det.;
ZMUC C-F-123118
. –
Vestjylland
(WJ)
•
Sneum Sluse
;
55°43.303′ N
,
8°60.710′ E
; MG74;
24 Mar. 2019
;
L. Kjaer-Thomsen
&
L. Thomas
1403
;
JP
det.;
ZMUC C-F-123949
.
On
Bembidion normannum
Dejean, 1831
(Col.
Carabidae
)
DENMARK
–
Fyn
(F)
•
Sprogø
;
55°20.012′ N
,
10°58.263′ E
; PG23;
7 Sep. 2013
;
JP
213
;
JP
det.;
ZMUC C-F-122692
.
On
Bembidion quadrimaculatum
(Linnaeus, 1761)
(Col.
Carabidae
)
DENMARK
–
Fyn
(F)
•
Gyldenstens
inddaemmede
Strand
;
55°34.447′ N
,
10°7.211′ E
; NG75;
2 Apr. 2014
;
JP
711
;
JP
det.;
ZMUC C-F-123201
.
On
Bembidion varium
(Olivier, 1795)
(Col.
Carabidae
)
DENMARK
–
Lolland, Falster, Møn
(LFM)
•
Ulvshale
;
55°3.218′ N
,
12°15.801′ E
; UB20;
10 Jun. 2017
;
JP
430
;
JP
det.;
ZMUC C-F-122915
.
On
Dyschirius globosus
(Herbst, 1784)
(Col.
Carabidae
)
DENMARK
–
Fyn
(F)
•
Gyldenstens
inddaemmede
Strand
;
55°34.447′ N
,
10°7.211′ E
; NG75;
2Apr. 2014
;
JP
693;
JP
det.;
ZMUC C-F-123181
. –
Lolland, Falster, Møn
(LFM)
•
Busemarke Mose
;
54°57.459′ N
,
12°27.054′ E
; UA39;
28 Aug. 2019
;
JP
1507
;
JP
det.;
ZMUC C-F-124094
•
Lysemose
ved
Maribo
;
54°46.971′ N
,
11°28.959′ E
; PF57;
4 Jun. 2018
;
JP
1127
;
JP
det.;
ZMUC C-F-123650
•
Vest
for
Bandholm
;
54°50.303′ N
,
11°28.050′ E
; PF57;
3 Jan. 2019
;
JP
1305
;
JP
det.;
ZMUC C-F-123840
. –
Nordøstjylland
(NEJ)
•
Vandplasken
ved
Kaersgård Strand
;
57°31.199′ N
,
9°53.070′ E
; NJ57;
21 Nov. 2017
;
JP
802
;
JP
det.;
ZMUC C-F-123297
. –
Nordøstsjaelland
(NEZ)
•
Gurre Vang
;
56°1.470′ N
,
12°29.486′ E
; UC41;
14 Apr. 2017
;
JP
269;
JP
det.;
ZMUC C-F-122748
•
Nord
for
Risø
;
55°42.184′ N
,
12°6.692′ E
; UB17;
14 Dec. 2013
;
R. Aagaard Jensen
623
;
JP
det.;
ZMUC C-F-123112
•
Ud
for
Kongelundsfortet
;
55°33.724′ N
,
12°33.731′ E
; UB46;
15 Apr. 2018
;
JP
1054
;
JP
det.;
ZMUC C-F-123570
. –
Nordvestjylland
(NWJ)
•
Syd
for
Stårup
;
56°34.792′ N
,
9°5.958′ E
; NH07;
6 Feb. 2015
;
JP
215
;
JP
det.;
ZMUC C-F-122695
•
Vest
for
Vinderup
;
56°28.727′ N
,
8°44.810′ E
; MH85;
19 Jan. 2015
;
JP
769
;
JP
det.;
ZMUC C-F-123264
. –
Nordvestsjaelland
(NWZ)
•
Flasken
nord for
Reersø
;
55°31.965′ N
,
11°9.831′ E
; PG35;
26 Jan. 2014
;
JP
610
;
JP
det.;
ZMUC C-F-123098
•
Flyndersø
ved
Dybesø
;
55°58.151′ N
,
11°45.621′ E
; PH60;
4 Nov. 2017
;
JP
713
;
JP
det.;
ZMUC C-F-123205
•
Nordbredden
af
Skarresø
;
55°39.425′ N
,
11°22.998′ E
; PG47;
1 May 2013
;
JP
854
;
JP
det.;
ZMUC C-F-123349
•
Nordøstbredden
af
Tissø
;
55°35.612′ N
,
11°18.461′ E
; PG46;
1 May 2013
;
JP
756;
JP
det.;
ZMUC C-F-123250
. –
Sydsjaelland
(SZ)
•
Even Bro
;
55°8.694′ N
,
12°0.601′ E
; UB11;
17 Feb. 2019
;
JP
1363
;
JP
det.;
ZMUC C-F-123901
•
Gammel Kalvehave
;
54°59.745′ N
,
12°8.232′ E
; UA19;
25 Apr. 2019
;
JP
1441
;
JP
det.;
ZMUC C-F-123989
•
Østerskov
ved
Langebaek
;
54°59.572′ N
,
12°6.715′ E
; UA19;
3 Jan. 2019
;
JP
1272
;
JP
det.;
ZMUC C-F-123802
•
Tryggevaelde Å ved
Varpelev
;
55°21.055′ N
,
12°15.877′ E
; UB23;
6 Jan. 2018
;
JP
962
;
JP
det.;
ZMUC C-F-123469
.
On
Dyschirius tristis
Stephens, 1827
(Col.
Carabidae
)
DENMARK
–
Nordøstsjaelland
(NEZ)
•
Eriksholm Skov
;
55°41.771′ N
,
11°47.912′ E
; PG77;
4 Nov. 2017
;
JP
743
;
JP
det.;
ZMUC C-F-123237
. –
Sydsjaelland
(SZ)
•
Even Bro
;
55°8.694′ N
,
12°0.601′ E
; UB11;
17 Feb. 2019
;
JP
1364
;
JP
det.;
ZMUC C-F-123903
.
Remarks
This is a very variable and may be a collective species in need of a thorough revision (
Haelewaters
et al.
2019b
). Attempts have been made to isolate some species with more or less luck (
Benjamin & Shanor 1952
;
Santamaria
et al.
2020a
). An example of this may be
L. tenera
which was described on
Bembidion dentellum
(Thunberg, 1787)
by
Majewski (1994b)
from
Poland
. It has been separated from
L. pedicellata
by several characters, especially related to the branching of outer appendage. We have found similar thalli to
L. tenera
on
Bembidion aeneum
(
Fig. 40C
). Distinguishing characteristics seem not to be very robust; furthermore, normal thalli of
L. pedicellata
have been found on the same host together with that
tenera
-like forms.
This species gives the name to a morphologically related group of species that share some characteristics such as: IV and V cells of the same height, and the presence of at least a first constricted and dark septum in the outer appendage.