Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species Author Yamasaki, Takeshi 0000-0002-2419-188X Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan & Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan & k 0468874 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2419 - 188 X k0468874@gmail.com Author Hashimoto, Yoshiaki 0000-0002-6696-5011 Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan & Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan & aenictus @ mac. com; https: // orcid. org / 0000 - 0002 - 6696 - 5011 aenictus@mac.com Author Endo, Tomoji 0009-0007-8279-9897 School of Human Sciences, Kobe College, Okadayama 4 - 1, Nishinomiya, Hyogo, 662 - 8505, Japan & endo @ mail. kobe-c. ac. jp; https: // orcid. org / 0009 - 0007 - 8279 - 9897 endo@mail.kobe-c.ac.jp Author Hyodo, Fujio 0000-0001-8344-9353 Research Core for Interdisciplinary Sciences, Okayama University, 3 - 1 - 1, Tsushimanaka, Okayama, 700 - 8530, Japan & fhyodo @ cc. okayama-u. ac. jp; https: // orcid. org / 0000 - 0001 - 8344 - 9353 fhyodo@cc.okayama-u.ac.jp Author Itioka, Takao 0000-0002-8295-5657 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida-nihonmatsu, Kyoto 606 - 8501, Japan & ichioka. takao. 5 m @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0002 - 8295 - 5657 ichioka.takao.5m@kyoto-u.ac.jp Author Mohamed, Maryati Tun Hussein Onn Malaysia, Hab Pendidikan Tinggi Pagoh, KM 1, Jalan Panchor, 84600 Panchor, Johor, Malaysia & maryatihayati 52 @ gmail. com Author Meleng, Paulus 0000-0003-3777-0597 Research Development and Innovation Division, Forest Department Sarawak, Kuching, KM 10, Jalan Datuk Amar Stephen Kalong Ningkan, 93250 Kuching, Sarawak, Malaysia & paulusm @ sarawak. gov. my; https: // orcid. org / 0000 - 0003 - 3777 - 0597 paulusm@sarawak.gov.my text Zootaxa 2023 2023-09-06 5343 1 55 73 http://dx.doi.org/10.11646/zootaxa.5343.1.3 journal article 10.11646/zootaxa.5343.1.3 1175-5326 8324250 20BF9AB4-F86D-42EA-AEAA-7C03EBA44EB6 Utivarachna ichneumon Deeleman-Reinhold, 2001 Figs 20–23 Utivarachna ichneumon Deeleman-Reinhold, 2001: 376 , figs 581–585. Type material examined. Holotype ♁ ( RMNH .ARA. 15223; Fig. 20E ), Danum Valley F. C. , E. Sabah , 6– 16. V .[1991], Coll. Deeleman , leg. C.L. & P. R . Deeleman. FIGURE 20. Utivarachna ichneumon Deeleman-Reinhold, 2001 , male, holotype (RNMH.ARA.15223). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars: A–C, 1 mm; D, 0.50 mm. FIGURE 21. Utivarachna ichneumon , male, holotype (RNMH.ARA.15223). A left palp, prolateral view; B same, ventral view; C same, retrolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum. Other material. 1 ♀ ( DV1 _20040910-E5), Danum Valley , Sabah , 10.IX.2004 , T . Endo leg.; 1 ♁ ( DV10 _ 20050222-I15), same locality, 22.II.2005 ; 1 ♀ ( DV7 _20050220-O16), same locality, 20.II.2005 , leg. R . Oie ; 1 ♀ ( DV53 _20080110-E7), same locality, 10.I.2008 , leg. T . Endo. All specimens are deposited at MNHAH . Diagnosis. See Diagnosis of U. fukasawana . Measurements ( holotype ♁/ ♀ ). Carapace length 3.95/3.16; cephalic width 1.50/1.37; thoracic width 1.93/1.80. Diameter of eyes: AME 0.18/0.17; ALE 0.16/0.15; PME 0.13/0.13; PLE 0.18/0.17. Interdistances between eyes: AME–AME 0.03/0.03; AME–ALE 0.07/0.07; ALE–ALE 0.56/0.51; PME–PME 0.21/0.20; PME–PLE 0.33/0.27; PLE–PLE 1.15/1.00; ALE–PLE 0.18/0.16; AME–PME 0.06/0.06. MOA: length 0.35/0.32; anterior width 0.40/0.37; posterior width 0.50/0.46. Clypeus height 0.32/0.36. Abdomen length 3.25/2.86; width 1.36/1.83. Legs: I 12.44 (3.35, 1.13, 3.33, 3.50, 1.13)/9.75 (2.73, 0.93, 2.46, 2.53, 1.10); II 10.00 (2.83, 0.82, 2.53, 2.66, 1.16)/8.31 (2.36, 0.73, 2.06, 2.16, 1.00); III 6.35 (1.77, 0.70, 1.42, 1.75, 0.71)/5.51 (1.52, 0.61, 1.20, 1.53, 0.65); IV 8.51 (2.37, 0.68, 2.00, 2.66, 0.80)/7.54 (2.07, 0.61, 1.75, 2.30, 0.81). Male ( Fig. 20A–D ). Carapace entirely granulated, dorsally oval; posterior stalk slightly shorter than 1/2 carapace length; PER recurved in dorsal view; ocular tubercle accompanied with PLE; thoracic lateral margins undulating. Chelicerae granulated, with five prolateral and five retrolateral teeth. Ventral clypeus margin with small wedge-shaped projection between chelicerae. Pedicel short. Abdomen elongate oval; anterior two-thirds covered with dorsal scutum, weakly developed. Leg I and II elongated, covered with spherical granulation; ventral sides of tibia, metatarsus and tarsus bearing cusps. Palp ( Fig. 21A–D ). Cymbium slender oval; length almost twice as wide. Bulb oval; posterior half of bulb weakly sclerotized with subtegulum, and anterior half membranous. Embolus coiled; basal part of embolus accompanied by slender sclerotized plate covered by membranous surface of bulb. Sperm duct running from retrolateral side of bulb, U-curved along posterior edge of bulb, then curving to prolateral side. RTA strongly extended, apically developed into semi-transparent plate with apical claw; prolateral corner of semi-transparent plate developed, and anterior margin transforming into short claw. Coloration and setation ( Fig. 20A–D ). Carapace and chelicerae brown, covered with fine setae. Abdomen cream, with pale brown dorsal scutum, entirely covered with small fine setae. All legs pale brown; leg I darker than others. Female ( Fig. 22A–D ). Somatic characters almost same as in males, except for abdomen shape and shorter posterior stalk; posterior stalk almost as long as 1/4 carapace length. Chelicerae granulated, with five or six prolateral and five retrolateral teeth. Abdomen oval, without dorsal scutum. Copulatory organs ( Fig. 23A–C ). Copulatory atrium longer than wide. Copulatory ducts curved into loop anteriorly, then intricately coiled. Bursae long, tube-like shaped, extending posteriorly. Spermathecae globular. Fertilization ducts straight, extending from spermathecae towards posterior. Coloration and setation ( Fig. 22A–D ). Almost same as in male. FIGURE 22. Utivarachna ichneumon Deeleman-Reinhold, 2001 , female (DV53). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view. Scale bars A–D: 1 mm. FIGURE 23. Utivarachna ichneumon , female (DV53). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca. Distribution. Sabah (known only from the type locality). Remarks. The non-type material examined in the present study was collected from the type locality, Danum Valley, Sabah state . The conspecificity of male and female specimens was based on the long stalk of carapace. Other Utivarachna species with long posterior stalks were not collected from the study site. The female is described here for the first time.