Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species
Author
Yamasaki, Takeshi
0000-0002-2419-188X
Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan & Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan & k 0468874 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2419 - 188 X
k0468874@gmail.com
Author
Hashimoto, Yoshiaki
0000-0002-6696-5011
Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan & Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan & aenictus @ mac. com; https: // orcid. org / 0000 - 0002 - 6696 - 5011
aenictus@mac.com
Author
Endo, Tomoji
0009-0007-8279-9897
School of Human Sciences, Kobe College, Okadayama 4 - 1, Nishinomiya, Hyogo, 662 - 8505, Japan & endo @ mail. kobe-c. ac. jp; https: // orcid. org / 0009 - 0007 - 8279 - 9897
endo@mail.kobe-c.ac.jp
Author
Hyodo, Fujio
0000-0001-8344-9353
Research Core for Interdisciplinary Sciences, Okayama University, 3 - 1 - 1, Tsushimanaka, Okayama, 700 - 8530, Japan & fhyodo @ cc. okayama-u. ac. jp; https: // orcid. org / 0000 - 0001 - 8344 - 9353
fhyodo@cc.okayama-u.ac.jp
Author
Itioka, Takao
0000-0002-8295-5657
Graduate School of Human and Environmental Studies, Kyoto University, Yoshida-nihonmatsu, Kyoto 606 - 8501, Japan & ichioka. takao. 5 m @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0002 - 8295 - 5657
ichioka.takao.5m@kyoto-u.ac.jp
Author
Mohamed, Maryati
Tun Hussein Onn Malaysia, Hab Pendidikan Tinggi Pagoh, KM 1, Jalan Panchor, 84600 Panchor, Johor, Malaysia & maryatihayati 52 @ gmail. com
Author
Meleng, Paulus
0000-0003-3777-0597
Research Development and Innovation Division, Forest Department Sarawak, Kuching, KM 10, Jalan Datuk Amar Stephen Kalong Ningkan, 93250 Kuching, Sarawak, Malaysia & paulusm @ sarawak. gov. my; https: // orcid. org / 0000 - 0003 - 3777 - 0597
paulusm@sarawak.gov.my
text
Zootaxa
2023
2023-09-06
5343
1
55
73
http://dx.doi.org/10.11646/zootaxa.5343.1.3
journal article
10.11646/zootaxa.5343.1.3
1175-5326
8324250
20BF9AB4-F86D-42EA-AEAA-7C03EBA44EB6
Utivarachna itiokai
Yamasaki
sp. nov.
Figs 1–3
Type material.
Holotype
♁
(
LMB
_
TY20180829
_TR01,
FRCS
),
Sungai Liku
,
Lambir Hills National Park
,
Sarawak
, BORNEO,
29.VIII.2018
, leg.
T
. Yamasaki.
Diagnosis.
Utivarachna itiokai
sp. nov.
matches the following characteristics of the
dusun
-group: the carapace is truncated at the posterior end; each of the posterior eyes is evenly spaced; the RTA is well-sclerotized apically without membranous parts; the legs I and II bear rounded cusps ventrally except for their trochanters and coxae; the proximal part of metatarsus I is concave ventrally.
The males of
U. itiokai
sp. nov.
are similar to those of
U. dusun
and
U. rubra
in the structures of the palp, but can be distinguished from the latter two species by the inclining orientation of the coils formed by the embolus: the plane of the embolus coils inclines retrolaterally and is covered by the cymbium in
U. itiokai
sp
.
nov
.
, whereas the plane of the coils inclines dorsally in
U. dusun
and
U. rubra
.
Etymology.
The specific name is given in honor of Dr. Takao Itioka (
Kyoto
University).
Measurements
(
holotype
♁). Carapace length 1.66; width 1.25. Cephalic width 0.90. Diameter of eyes: AME 0.10; ALE 0.11; PME 0.11; PLE 0.11. Interdistances between eyes: AME–AME 0.03; AME–ALE 0.03; ALE–ALE 0.30; PME–PME 0.10; PME–PLE 0.12; PLE–PLE 0.56; ALE–PLE 0.11; AME–PME 0.05. MOA: length 0.25; anterior width 0.25; posterior width 0.31. Clypeus height 0.18. Abdomen length 1.77; width 1.25. Legs: I 4.32 (1.38, 0.61, 1.06, 0.71, 0.56); II 3.47 (1.00, 0.45, 0.76, 0.76, 0.50); III 2.67 (0.78, 0.37, 0.52, 0.65, 0.35); IV 3.32 (0.91, 0.40, 0.75, 0.86, 0.40).
Male
(
Fig. 1A–E
). Carapace oval and superficially granulated, with truncated posterior end. Thoracic lateral margins hardly undulating. PER longer than AER, slightly recurved. Fovea shallow, indistinct. Ventral clypeus margin with small truncated projection between chelicerae. Chelicerae superficially granulated, with three teeth on each prolateral and retrolateral margin. Pedicel short. Abdomen oval; entire dorsum covered by sclerotized scutum. Leg I enlarged, with ventral cusps; Leg II with ventral cusps. Leg III and IV with sparse ventral tubercles on tibiae.
FIGURE 1.
Utivarachna itiokai
sp. nov.
, male, holotype (LMB_TY20180829_TR01).
A
habitus, dorsal view;
B
same, ventral view;
C
same, lateral view;
D
cephalic part and chelicerae, frontal view;
E
left leg I, prolateral view. Scale bars: 1 mm.
FIGURE 2.
Utivarachna itiokai
sp. nov.
, male, holotype (LMB_TY20180829_TR01).
A
left palp, prolateral view;
B
same, ventral view;
C
same, retrolateral view;
D
same, dorsal view. Scale bars: 0.50 mm. E—embolus, St—subtegulum.
FIGURE 3.
Utivarachna itiokai
sp. nov.
, male, holotype (LMB_TY20180829_TR01).
A
left palp, prolateral view;
B
same, ventral view;
C
same, retrolateral view. Scale bars: 0.25 mm. E—embolus, St—subtegulum
Palp (
Figs 2A–D
,
3A–C
). Cymbium oval. Bulb oval. Sperm duct U-shaped, visible through median tegulum. Embolus coiled in horizontal plane, coils inclined towards retrolateral side of cymbium, partially covered by cymbium; embolar base sclerotized, forming median plate, originated in middle of tegulum. RTA long in retrolateral view, shorter than half of cymbium; apically strongly dorsally curved, apex not reaching dorsal cymbium outline.
Coloration and setation (
Fig. 1A–E
). Carapace and chelicera brown. Abdomen covered with fine setae; dorsal scutum gray; lateral and ventral surfaces cream, tinged with black.
Female.
Unknown.
Distribution.
Sarawak
(known only from the
type
locality).