3584 Author Michat, Mariano C. Author Alarie, Yves Author Watts, Chris H. S. text Zootaxa 2012 2012-12-13 3584 1 1 110 http://dx.doi.org/10.11646/zootaxa.3584.1.1 journal article 10.11646/zootaxa.3584.1.1 1175­5334 10096816 072032C4-63FC-499A-A61D-58B428051302 Limbodessus raesideensis (Watts & Humphreys, 2001) ( Figs 212–215 ) Source of material. Two specimens of instar II and two of instar III were used for the description ( Table 1 ). Larvae were collected in association with adults at the following localities: Australia , Raeside palaeovalley , Lake Mason calcrete, Salt Well , BES 8353, 27.54S , 119.62417E , 24–VI–2000 , coll. W. F. Humphreys and S. Hinze ; Lake Mason calcrete, Salt Well , BES 14361, 27.586S , 119.5218E , 20–IX–2006 , coll. W. F. Humphreys and T. Moulds. Diagnosis (instar III). Larger species (HL more than 1.25 mm ); head ( Fig. 212 ) pyriform; nasale digitiform; half-circle of dense spinulae on lateroventral margins of nasale absent; hole-like structure on ventrodistal surface of nasale absent; lateral margins of nasale not inflated in dorsal view; lateral branches of nasale well developed; slender spinulae anterior to seta FR13 numerous (25 or more); occipital foramen strongly reduced (HW/OCW TABLE 18. Number and position of secondary setae on the legs of third-instar larvae of species of Limbodessus . Numbers between slash marks refer to pro-, meso- and metathoracic leg, respectively. A = anterior, D = dorsal, P = posterior, Pr = proximal, V = ventral, Total = total number of secondary setae on the segment (excluding primary setae).

Segment	Position	L. inornatus	L. leysi	L. masonensis	L. millbilliensis
Coxa	A	10–13 / 10–18 / 17–26	0 / 0 / 0	0 / 0 / 0	0 / 0 / 0
P	3–7 / 14–16 / 18–22	0 / 0 / 0	0 / 0 / 0	0 / 0 / 0
PD	7 / 4–6 / 5–8	2 / 2–3 / 1–2	3–5 / 3–4 / 1–3	3 / 3 / 2
V	9–11 / 9–10 / 7–11	1–2 / 1 / 1	2 / 1 / 1–2	1 / 1 / 1
Total	32–35 / 40–47 / 56–58	3–4 / 3–4 / 2–3	5–7 / 4–5 / 2–5	4 / 4 / 3
Trochanter	Pr	0 / 1 / 1–2	0–1 / 1 / 1	0 / 1 / 1	0 / 1 / 1
Total	0 / 1 / 1–2	0–1 / 1 / 1	0 / 1 / 1	0 / 1 / 1
Femur	AD	2 / 2–3 / 3–4	2 / 2–3 / 2	2 / 2 / 2–3	2 / 2 / 2
AV	8–9 / 9 / 9–11	3–4 / 1–4 / 3–4	3–5 / 4 / 4–5	3–4 / 3–4 / 4–5
PD	0 / 0 / 0	0 / 0 / 0	0 / 0 / 0	0 / 0 / 0
PV	5 / 7–9 / 10–13	1–3 / 2–3 / 4	4 / 5 / 4–5	3 / 3 / 4–5
Total	15–16 / 18–21 / 23–27	6–9 / 7–9 / 9–10	9–11 / 11 / 11–12	8–9 / 8–9 / 10–12
Tibia	AD	0 / 1 / 3	0 / 1 / 1	1 / 1–2 / 1	1 / 1 / 1
AV	0 / 2–3 / 2–3	1 / 1–2 / 1–3	0 / 2 / 2–3	0 / 2 / 2–3
PD	1 / 2 / 2–3	0–1 / 1 / 1	0 / 1 / 1	0 / 1 / 1
PV	1–2 / 2–3 / 4–6	0–1 / 1–2 / 0–2	1 / 1 / 1	1 / 1 / 1–2
Total	2–3 / 7–9 / 11–15	1–3 / 5 / 5	2 / 5–6 / 5–6	2 / 5 / 5–7
Tarsus	AD	0 / 0 / 0	0 / 0 / 0	0 / 0 / 0	0 / 0 / 0
AV	0 / 2 / 3–4	1 / 2 / 2	0 / 1 / 1–2	0 / 2 / 2–3
PD	0 / 2 / 2–3	0 / 1 / 1	0 / 1 / 1	0 / 1 / 1
PV	1 / 2 / 4	0 / 0 / 0	0 / 0 / 0	0 / 0 / 0
Total	1 / 6 / 9–11	1 / 3 / 3	0 / 2 / 2–3	0 / 3 / 3–4

Instar I. Not available. Instar II. Head. A3 less than 2.50 times longer than A1; A4 less than 0.70 times as long as A3; MN less than 4.70 times longer than broad; MP2 less than 1.05 times longer than MP1; MP2 more than 3.20 times longer than MP3; LP2 less than 0.90 times as long as LP1. Legs. L3 2.25–2.95 times longer than HW; CL(L3) less than 0.40 times as long as TA. Abdomen. U 3.00–4.00 times longer than LAS; U less than 2.15 times longer than HW; U1 more than 1.65 times longer than U2. Chaetotaxy. Anteroventral margin of nasale with 70 lamellae clypeales distributed in 1–2 rows; anterior secondary setae on proCO absent; meso- and metaCO with less than 5 posterodorsal secondary setae; ventral secondary setae on pro- and mesoCO present; proFE with more than 3 posteroventral secondary setae; metaFE with more than 11 secondary setae; anterodorsal secondary setae on proTI absent; anteroventral, posterodorsal and posteroventral secondary setae on proTI present; anterodorsal and posteroventral secondary setae on meso- and metaTI present; meso- and metaTI with less than 2 posterodorsal secondary setae; metaTI with more than 5 secondary setae; anterodorsal secondary setae on proTA absent; anteroventral and posteroventral secondary setae on proTA present; posterodorsal and posteroventral secondary setae on meso- and metaTA present; metaTA with more than 5 secondary setae; secondary setae on U present. Measurements and ratios that characterize the body shape are shown in Table 7 . Secondary leg setation detailed in Table 15 . Instar III ( Figs 212–215 ). Head ( Fig. 212 ). A3 less than 2.25 times longer than A1; A3 less than 1.45 times longer than A2; MN less than 4.85 times longer than broad; MP more than 1.75 times longer than labial palpus; MP2 3.20–4.00 times longer than MP3; LP2 more than 0.65 times as long as LP1. Legs ( Figs 213–214 ). L3 2.25–2.85 times longer than HW. Abdomen ( Fig. 215 ). U more than 3.10 times longer than LAS; U 1.55–2.25 times longer than HW; U1 1.80–2.60 times longer than U2. Chaetotaxy. Anteroventral margin of nasale with 134 lamellae clypeales distributed in 2 rows; proCO with more than 7 anterior secondary setae; mesoCO with 1–5 anterior secondary setae; pro-, meso- and metaCO without posterior secondary setae; proCO with more than 21 secondary setae; metaCO with less than 20 secondary setae; anterodorsal secondary setae on pro- and metaFE present; metaFE with more than 12 anteroventral secondary setae; posterodorsal secondary setae on meso- and metaFE present; metaFE with more than 30 secondary setae; anterodorsal secondary setae on proTI absent; anteroventral and posterodorsal secondary setae on proTI present; mesoTI with more than 4 anteroventral secondary setae; metaTI with less than 9 anteroventral secondary setae; metaTI with more than 18 secondary setae; anterodorsal secondary setae on proTA absent; anterodorsal secondary setae on meso- and metaTA present; anteroventral and posterodorsal secondary setae on pro-, meso- and metaTA present; posteroventral secondary setae on pro- and mesoTA present; metaTA with 1–5 posteroventral secondary setae; proTA with 1–7 secondary setae; mesoTA with more than 7 secondary setae; metaTA with less than 16 secondary setae; secondary setae on U present. Measurements and ratios that characterize the body shape are shown in Table 11 . Secondary leg setation detailed in Table 19 . Remarks. Limbodessus raesideensis is a very characteristic stygobitic species that belongs to the group of species characterized by the presence of secondary setae on the urogomphus ( L. barwidgeeensis , L. bigbellensis , L. cooperi , L. eberhardi , L. macrohinkleri , L. nambiensis , L. yandalensis ) ( Fig. 215 ). Within this group, it can be distinguished by the following combination of characters: head pyriform ( Fig. 212 ), occipital suture present ( Fig. 212 ), occipital foramen strongly reduced ( Fig. 212 ), presence of anterior secondary setae on the coxa ( Fig. 213 ), presence of posterodorsal secondary setae on the meso- and metafemur, femur with more than 30 secondary setae, and absence of anterodorsal secondary setae on the protibia and protarsus ( Fig. 213 ).